The Scale April 2002

Editors:
Dug Miller & Gary Miller
Systematic Entomol. Lab.
Plant Sciences Institute
Agric. Res. Service, USDA
Building 005, Room 137
BARC-West, Beltsville, MD
20705-2350 USA
FAX 01 301-504-6482
dmiller@sel.barc.usda.gov

Issues from 1996 to the present can be downloaded from the following web page: http://www.sel.barc.usda.gov/coccoidea/scaleframe.html

The meeting in Padua, Italy was extraordinary, to say the least. The coccidology community wants to again extend its congratulations and thanks to Giuseppina Pellizzari and her many contributing colleagues for making this meeting so very special. Photographs of the meeting can be viewed at: http://www.sel.barc.usda.gov/Coccoidea/Padua/destination/FrameSet.htm

We are pleased to announce that the X International Symposium on Scale Insect Studies will be held in Adana, Turkey in April 2004. This city is one of the largest in Turkey and is located along the eastern Mediterranean coast. The Tauros Mountains are situated to the north of the city and a typical Mediterranean Coast line is present to the south. Nearby Cukurove is present between the Seyhan and Ceyhan rivers and it contains some of the most fertile areas in Turkey. It is famous for intensive agricultural production including mainly field crops, maize, cotton, fruit crops, and citrus. In ancient times this area called Kilikya and was of special interest because of its rich soils. April is a great time to visit Adana with the sweet aroma of citrus blossoms permeating the air and mild spring weather. There are four or five flights each day from Istanbul which will make travel plans easy since the Adana airport is close to the meeting site. The symposium will discuss all aspects of Coccidiology including: Systematics, Biology, Scale insect pests, Zoogeography, Biological control, Monitoring. A web page is currently being established.

This web site continues to flourish. The address is: http://193.204.185.103/scaleinfo/ scale.htm. The most recent table of contents is as follows:

Do we need to change the rank of the higher taxa of scale insects (Hemiptera: Sternorrhyncha: Coccoidea)? By Penny Gullan

On the term describing the post-embryonic development of Diaspididae (Rhynchota Coccoidea). by Francesco Porcelli

Editor’s Note: If you have news that you would like to have included in the next edition of the Scale, we would be happy to receive it. E-mails are especially appreciated.

Samir A. El-Serwy, Plant Protection Research Institute, Egypt: The following are abstracts of papers that were given in 2001: El- Serwy, S. (2001). Ecology, biology and natural enemies of the red-striped soft scale, Pulvinaria tenuivalvata (Newstead) ( Hemiptera: Coccidae), a pest of sugarcane in Egypt. [ Abstract of presentation at the IX ISSIS, Sep. 2- 8, 2001 in Padua, Italy.]

And El- Serwy, S. (2001). Parasitic complex of Pulvinaria tenuivalvata (Newstead) in Egypt. [Abstract at the ESA Annual Meeting, Dec. 9- 12, 2001 in San Diego, California, USA.]

Penny Gullan, University of California, USA: I've now spent two years living in the USA and working at the University of California in Davis (UCD). 2001 was a busy and fairly productive period. I taught in two new undergraduate courses, one on systematic entomology and the other on biodiversity, and worked on a collaborative project on mealybug bacterial endosymbionts with Drs Paul Baumann and My-Lo Thao of the Microbiology Department at UCD. Paul and My-Lo have sequenced genes from the primary and secondary endosymbionts of a number of mealybug species that I collected/acquired and identified. We recently completed a manuscript on endosymbiont coevolution. A new Ph.D. student, Nam Nguyen, started his studies with me in September 2001. Nam will be working on mealybug phylogenetic systematics using morphological data, but presently is completing lots of coursework. During May, Doug Williams and Dug Miller visited UCD for the last of three visits funded by an NSF Curation grant to the Director of the Bohart Museum of Entomology, Dr. Lynn Kimsey. Doug and Dug continued to assist with sorting of the Ferris-McKenzie scale insect collection, especially identification of slide-mounts and the updating of names of armored scales. At the same time, Ben Normark visited Penny's lab for a week and studied armored scales, did some collecting and presented a seminar on "Phylogeny and alternative genetic systems in scale insects". Ray Gill came to UCD to assist in the coccidological event whenever he could escape administration at the California Department of Food and Agriculture. I had a thoroughly enjoyable time with Doug, Dug, Ben and occasionally Ray in my lab, and Bohart Museum scale insect collection benefited greatly too. Another highlight of the year was spending time in Italy to attend the IXth ISSIS in Padua and then to have a short holiday in NE Italy. Peter Cranston, Nam and I all had a wonderful visit and much appreciated the efficient organization and friendly hospitality of Giuseppina Pellizzari and her colleagues and students. The third highlight of the year was the award of a 5-year National Science Foundation (NSF) grant as part of the PEET Program (Partnerships for Enhancing Expertise in Taxonomy) to conduct research and training in phylogenetic systematics of scale insects. For more information, check out: http://www.nsf.gov/ nsf /nsfpubs/nsf9721.htm. The project is centered at UCD but involves active collaboration with Dug Miller and Ben Normark who will assist me with the training of two graduate students, several undergraduate students and a postdoctoral researcher. The proposed postdoctoral researcher is already a scale insect fanatic: Takumasa Kondo (Demian to his friends) will need little introduction to those who were at the last ISSIS. Currently Demian is completing a Ph.D. on the New World Myzolecaniinae with Mike Williams at Auburn University and hopes to move to Davis later in 2002 to study SE Asian and Australasian Myzolecaniinae. In late December 2001 and early January 2002, I did fieldwork in southern Africa in conjunction with the field studies of Peter Cranston. Scale insect collecting was fairly sparse but I did find a few interesting things, especially some gall-inducing Grewiacoccus mealybugs on a small Grewia tree in Zimbabwe, and other mealybugs on the bizarre desert-living gymnosperm Welwitschia mirabilis. From January 15-18, 2002, I attended the Third International Symposium on "The Biology of Gall-Inducing Arthropods", held at the University of Stellenbosch, South Africa (organized by Schalk Louw, Jan Giliomee and their collegaues). There I presented the only talk on scale insects in collaboration with Jan Giliomee and Chris Hodgson on the South African gall-inducing Calycicoccus merwei (Eriococcidae), and a poster on Apiomorpha with Lyn Cook. During the symposium I was shocked and very saddened to learn of the death of our dear colleague Salvatore Marotta. From Africa I travelled on to Canberra, Australia, where I collaborated with Lyn Cook in my former department of Botany & Zoology at the Australian National University. Lyn and I worked on two manuscripts: one is a book chapter on gall-inducing scale insects (also co-authored with Dug Miller) and the other is a paper on the relationships of gall-inducing eriococcid scales as estimated from molecular data. When my departure day of February 6^th arrived I was reluctant to leave Canberra and return to the end of a Californian winter (although Californian winters can hardly be called winters). Since my return, I've been kept very busy trying to complete my parts of that chapter on gall-inducing coccoids and recently organizing a symposium on the Phylogeny and Evolution of Sternorrhyncha for the next Annual Entomological Society of America (ESA) meeting to be held at Fort Lauderdale in Florida in November this year. See http://www.entsoc.org/annual_meeting/2002 for details of the Annual ESA meeting, although details of the Sternorrhyncha symposium are not yet available. There's one thing about which you can be certain concerning the symposium -- the Sternorrhyncha will be a treated as a suborder of the order Hemiptera!

In January 2002, specimens of a second eastern hemisphere species of the same genus, A. tubercularis Newstead, known as the white mango scale, was discovered on a mango in the Miami area. Surveys have located additional specimens in nearby areas. Thus, it is considered established in Florida. The lac insects, family Kerriidae, are native to the eastern hemisphere, and recently, Paratachardina lobata lobata (Chamberlin) was found in southern Florida. This insect was first detected here in 1999 on a single Hibiscus rosa-sinensis. The plant was destroyed in an attempt to eradicate the insect. However, the scale was again found in March 2000, and is currently reported throughout southeastern Florida. In this region, Bucida buceras, Chrysobalanus icaco, Conocarpus erectus, Ficus spp., Hibiscus, Myrica cerifera, and Quercus geminata are frequent hosts of this insect, but data on its host relations is preliminary. We are currently planning studies of the biology and control of this insect. In November 2001, the Diaspididae, Duplachionaspis divergens (Green) was discovered in widely separated areas of Florida. Future plans for this scale are still under review and evaluation. The known hosts are all in the Poaceae. Reference: Howard, FW., A. Hamon, T. M. McLaughliln, T. Weissling, and Si-Lin Yang. 1999. Aulacaspis yasumatsui (Hemiptera: Sternorrhyncha: Diaspididae), a scale insect pest of cycads recently introduced into Florida. Florida Entomologist 82 (1): 14-27

Michael Kosztarab, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, USA. The following are the highlights of my activities for the past two years. I was fortunate to have had two very busy but fruitful retirement years. In June 2000 we visited Evelyna Danzig in the beautiful Russian city of St. Petersburg. After seeing her workplace in the Zoological Institute Matilda, and I were taken to her lovely home for a tea party. It gave me much pleasure to see my article on the “Status and Future of Human Resources in Coccidology” finally printed in Entomologica (Bari) 33:19-34. During 2001 I also served as coeditor of the 380 page book: A History of the Virginia Academy of Science 1923-2001. I have reviewed a National Science Foundation grant proposal for coccidological research and a number of manuscripts for the ISSIS-IX event in Padua, Italy; and assisted the organizers for a few days, made a presentation and lead discussions at one of the sessions. Last December I attended the Entomological Society of America (ESA) Annual meeting and the Entomological Collections Network sessions in San Diego, California. There were several oral and poster presentations given on research with scale insects. Besides Ray Gill and Mike Williams, two Russian chalcid wasp experts, S.N. Myartseva ad V.A. Triapitsyn also attended the meetings. Apparently there were sufficient papers/posters on scales and other Sternorrhyncha research to qualify for an informal or formal session. Therefore, I was pleased to learn recently that a formal Sternorrhyncha Section A Symposium is planned for the November 2002 ESA national meeting in Fort Lauderdale, Florida by Penny Gullan. It is sad for a professor to learn that some of his former graduate students are reaching retirement, as it was with James O. Howell last year and will be with Avas B. Hamon this year. Last December we were all saddened to learn about the passing away, after a prolonged illness, our colleague and dear friend Salvatore Marotta. On the bright side I have learned that Dug Miller is the recipient of the L. O. Howard Lifetime Achievement Award at the Eastern Branch of ESA in 2002. Congratulations to Dug from our worldwide coccidologist community for receiving this well-earned recognition! It looks good for coccidology, especially since he is the second scale insect specialist in two years to be honored with this award. [editor’s note; Michael received the award for 2001]. Some of you may also recall that in 2000 the ESA President’s Award for a High School teacher successfully using insects in a class project was given for a cochineal scale project to Ms. Michelle Leddell of Santa Monica, California. So, national recognition for coccidological work has been received for three consecutive years. Let’s keep up with this good start!

Ferenc Kozár, Plant Protection Institute, Budapest, Hungary: I am sorry to report that I have not yet had the opportunity to finish the monograph on the Ortheziidae. However, it is near completion and will be in press soon. Because of my obligation to support my research program with grant funds, I have begun working on a project on the mealybug genus Rhizoecus. Based on extensive samples collected in Berlese samples from many parts of the world, I have discovered about 15 new species of the genus. Surprisingly, some of them are quite interesting. I am currently studying the types and unidentified material from several of the major museums. In fact, this note is written in Beltsville, Maryland where I am studying the extensive Rhizoecus material in the collection of the U.S. National Museum of Natural History. I also am enjoying my stay in the Miller Hotel. If you have material of Rhizoecus that you think should be included in this study, please contact me as soon as possible.

Ian Millar, National Collection of Insects, Pretoria, South Africa: I have been collaborating with Chris Hodgson (National Museum of Wales, Cardiff, U.K.) on a study of several interesting new species of South African Aclerdidae. This project has resulted in the description of a new subfamily, two new genera and three new species from southern Africa, and a phylogenetic analysis of the Aclerdidae. The work will be published, with Chris as senior author, in Systematic Entomology (the final manuscript was accepted in January this year for publication). In another project, I have compiled a key to the 50 genera of mealybugs that are known to occur in South Africa. The manuscript was reviewed last year, and is currently awaiting publication in African Entomology, the journal of the Entomological Society of Southern Africa. Lastly, I am editing a computer database of specimen information associated with the 6700 Coccoidea accessions in our collection. The database will enable searching of the collection's data by category, such as locality or host plant.

R. K. Varshney, Aligarh, India: I would like to draw attention to a relatively new journal that I have started. It is called Bionotes and is a quarterly newsletter that is in its fourth year. It is an outlet for short notes not exceeding four pages.

Douglas J. Williams, The Natural History Museum, Britian: Continues toward his goal of finishing his book on the mealybugs of southern Asia. The illustrations are nearly complete (just a few more Pseudococcus to do) and he is more that 3/4 of the way through the descriptions. He did the usual jaunt to Beltsville, Maryland and Davis, California in the spring of 2001 and is especially grateful to Penny Gullan and Peter Cranston for allowing him to stay in their beautiful house in the foothills during his visit there. He also completed a small paper with Dug Miller on the mealybug genus Antonina. He has numerous other projects either in press or near completion, but it is hard to keep track of all of them because there are so many.

It is particularly sad to announce the death of Salvatore Marotta. The short announcement from Giuseppina says it all. I am very sad to inform you that our dear friend and colleague Salvatore Marotta passed away on December 18, 2001. Those who were privileged to know him were very much aware of his generous nature, his kindness and goodness of heart, and his high level of scientific achievement and productivity. A quick analysis of his coccidology career as compiled by ScaleNet shows that he published 43 papers with the first appearing in 1983. His interests were highly diverse including invasive species, life history, morphology, nomenclature, systematics, and zoogeography to name a few. He was interested in all groups of scale insects but published papers that focused on Coccidae, Dactylopiidae, Diaspididae, Eriococcidae, Kermesidae, Margarodidae, Micrococcidae, Pseudococcidae, and Putoidae. Salvatore was a delightful human being who had a wonderful sense of humor. His enthusiasm and excitement for scale insect research was truly contagious. You couldn’t talk with him for even a few minutes without being struck by his intense interest in scale insects. He will be GREATLY MISSED, even by those of us who live on the other side of the world.

During the previous 12 months I've entered 1757 new records. The total in ScaleNet as of the last upgrade was 16,227. (Last year at this time I had entered 1283 records, and the year before, 790.) We might want to remind people that many of these newly entered titles are older ones that are just being discovered. Anyone who wants to keep track retrospectively of a certain topic or taxa may want to repeat their ScaleNet searches periodically, and not limit their searches to recently published papers.

Abd El-Salam, A. & Mangoud, H. 2001. Development and implementation of integrated pest management to programs of apple trees in reclaimed lands in Egypt: I - The fig scale insect (FSI), Russellaspis (Asterolecanium) pustulans (Cockerell). Journal of Agriculture in the Tropics and Subtropics 102(1):33-44. [AbdElSMa2001]

Notes: The fig scale insect (FSI), Russellaspis (Asterolecanium) pustulans (Cockerell) (Homoptera: Asterolecaniidae) is a serious pest of apple trees, Pyrus malus (Mill), in new reclaimed lands in Egypt. Both nymphs and adult females were found on apple trees all over the year. The distribution studies of the FSI on various parts of apple trees were carried out in a private farm located in El-Mansoria Center at El-Giza Governorate in Egypt. The distribution studies are equally important, particularly when oriented spraying is practiced. The data showed that 51,6 and 46.6% of adults and nymphs stages respectively of the FSI were concentrated on trunks, while, 48.4 and 53.4% respectively were distributed between main branches, new branches and new leaf petioles. The same trend was observed with the distribution of parasites. The FSI population was concentrated in the direction of water pipe in new reclaimed lands. Two methods were carried out to control this pest: the whole tree spraying and the oriented spraying. In the whole tree spraying, the whole tree was sprayed; about 7 liters of pesticide solution at winter time (after pruning) and 15 liters at summer time (before pruning). In the oriented spraying (spraying of infested branches), about 2.5 liter at winter time (after pruning) and 6 liters at summer time (before pruning) spraying solution per tree were sufficient to cover the infested branches. Both spraying methods (oriented and whole tree spraying) gave complete protection for 5 months (in winter) and 4 months (in summer) after which, reinfestation was observed. The whole tree spraying was the more expensive variants (235 and 162 LE/feddan) while the oriented spraying variants (106 and 71 LE/feddan) was relatively safe to the environment and also gave good reduction against the FSI pest and is very important in Integrated Pest Management (IPM) methods.

Abd Rabou, S. 2000. Parasitoids attacking Saccharicoccus sacchari (Cockerell) (Hemiptera: Pseudococcidae) on sugarcane in Egypt. 72-75. In: Allsopp, P.G. (Ed.) & Suasa Ard, W., Sugarcane Pest Management in the New Millennium. 4th Sugarcane Entomology Workshop International Society of Sugar Cane International Society of Sugar Cane Technologists, Indooroopilly, Australia. 104 pp. [AbdRab2000]

Notes: Sugarcane mealybug, Saccharicoccus sacchari (Cockerell), exerts drastic effects on sugarcane plants and is considered one of the most important pests attacking this plant in Egypt. The present work includes a survey of the parasitoids of this pest and their abundance in locations affected by it. The survey of S. sacchari parasitoids and hyperparasitoids took place during June 1998-June 1999. Nine parasitoids and hyperparasitoids were recorded. These are Anagyrus greeni Howard, A. pseudococci (Girault), Chartocerus subaenus (Foerster), Leptomastidea abnormis (Girault), Microterys sp., Paraphaenaodiscus sp., Prochiloneurus sp., Rhopus nigriclavus (Girault) and Rhopus sp. Abundance of these species was evaluated at three localities in Egypt. L. abnormis was the dominant parasitoid, showing maximum parasitism rates of 14 and 21% during October 1998 in Assiut and Qena governorates, respectively, while at the third locality it was not active. From these results, it is clear that there is an urgent need for introduction of other effective parasitoids from abroad to achieve effective control of this pest.

Abd-Rabou, S. 2001 (1999). Parasitoids attacking the Mediterranean black scale, Saissetia oleae (Hemiptera: Coccidae) on olive in Egypt. Entomologica 33 (1999): 169-172. [AbdRab2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Saissetia oleae is an important pest of olive trees in Egypt. A survey of the parasitoids of S. oleae was carried out monthly between April 1995 and March 1997 in three different locations in Egypt. Five species of Encyrtidae, a species of Pteromalidae and an aphelinid hyperparasite were found. Three of these records were new for Egypt.

Abd-Rabou, S. 2001a (1999). An annotated list of the Hymenopterous parasitoids of the Diaspididae (Hemiptera: coccoidea) in Egypt, with new records. Entomologica 33: 173-177. [AbdRab2001a]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Eighteen species of hymenopterous parasitoid of armoured scale insects (Hemiptera: Diaspididae) were recorded in a survey of host plants in three locations in Egypt during 1994-1997. The 16 species of Aphelinidae and two Encyrtidae are listed, along with their diaspidid hosts and location in Egypt; ten species were new records for Egypt.

Abd-Rabou, S., Hanafi, A. & Hussein, N. 2001 (1999). Notes on the parasitoids of the soft brown scale, Coccus hesperidum (Hemiptera: Coccidae) in Egypt. Entomologica 33(1999): 179-184. [AbdRabHaHu2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Thirteen species of hymenopterous parasitoids were found in association with Coccus hesperidum at three locations in Egypt: 4 species of Aphelinidae (of which three were recorded for the first time); seven species of Encyrtidae (of which three were recorded for the first time), the first record of a species of Mymaridae, plus a species of Pteromalidae. The relative abundance of these parasitoids is discussed.

Abdel-Salam, A.H. & Abdel-Baky, N.F. 2001. Life table and biological studies of Harmonia axyridis Pallas (Col., Coccinellidae) reared on the grain moth eggs of Sitotroga cerealella Olivier (Lep., Gelechiidae). Journal of Applied Entomology 125(8): 455-462. [AbdelSAb2001]

Notes: The developmental period of immature stages, survival per cent, longevity, fecundity and life table parameters of Harmonia axyridis by feeding on fresh and frozen grain moth eggs (GME) of Sitotroga cerealella were studied under laboratory conditions. The effect of crowding adults on fecundity of females was also determined. The total developmental time from egg hatching to adult eclosion ranged from 18.89 +/- 0.32 to 22.5 +/- 0.21 days on fresh and frozen GME, respectively. Survival per cent from egg hatching 0o adult emergence differed significantly when the predator fed on the two diets of GME. Also, pupal and adult weights were affected by feeding on fresh and frozen GME. There were no significant differences in longevity of females, while there were significant variations in fecundity of females and longevity of males. The calculated values of T, DT, R-o, r(m), and e(rm) were high by feeding on fresh GME. Moreover, rates of survivorship (Lx), and maximum oviposition per female per day (Mx) were higher when the predator was reared on fresh than on frozen GME. Crowding conditions of predator adults affected the fecundity of females due to egg cannibalism by both males and females of the predator. [Keyword: Margarodidae]

Abdrashitova, N. 2001 (1999). Dendrophilous Coccoids (Hemiptera: Coccoidea) of the walnut-fruit forest area of Kyrgyzstan. Entomologica 33 (1999): 229. [Abdras2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. A study of dendrophilous coccoid fauna of the walnut-fruit forest area of southern Kyrgyzstan (the Fergana and Chatkal mountain ranges) was undertaken. Forty-five coccoid species were discovered belonging to 27 genera and 6 families: Diaspididae (9 genera, 18 species), Coccidae (9 genera, 15 species), Pseudococcidae (6 genera, 9 species) and Margarodidae, Eriococcidae and Ortheziidae (1 species each). The scale insect fauna contains 9 species that are basically from Central Asia (20%) and 14 species which are subendemic to the Iranian and Turanian regions (31%). The rest of the species are Mediterranean (7 spp., 16%), European (2 spp., 5%), Palaearctic (4 spp., 9%) and Holarctic (3 spp., 7%). The complex of accidental species includes 6 species (13.5%). These scale species could be referred to three groups: polyphagous - 24 species (53.3%), oligophagous - 18 species (40%) and monophagous - 3 species (6.7%). In addition, on the basis of their frequency and damage, 10 species were categorised as numerous, 21 species as moderately numerous and 14 species as rare or infrequent. The most numerous and economically important species was Sphaerolecanium prunastri (Fonsc.), which caused extensive damage to the local alycha species (Prunus sogdiana Vass.). In these areas of Kyrgyzstan, biological control of the scale insect fauna is considered the most appropriate and promising method and such encyrtid parasitoids as Discodes coccophagus Ratz. and Microterys hortulanus Erd. play an important role in the control of S. prunastri.

Abdul, N.H. 2000. [Saissetia oleae (Bernard), occasional scale of Lebanon cedar (Hemiptera, Coccoidea, Lecanidae).] Saissetia oleae (Bernard), cochenille occasionnelle du Cedre au Liban (Hemiptera, Coccoidea, Lecanidae). (In French). Nouvelle Revue d'Entomologie 17(1): 81-82. [Abdul2000]

Notes: Individuals of Saissetia oleae were found on Lebanon cedar (Cedrus libani) in the region of Jbeil, Byblos, at 500 m altitude.

Abou-Elkhair, S. 2001 (1999). Scale insects (Hemiptera: Coccoidea) and their parasitoids on ornamental plants in Alexandria, Egypt. Entomologica 33 (1999): 185-195. [AbouEl2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] This paper discusses the scale insects (Coccoidea) and their parasitoids on ornamental plants in the Alexandria district of Egypt, between 1995 and 1998. Thirty-nine coccoid species belonging to eight families were noted: Asterolecaniidae, Coccidae, Diaspididae, Eriococcidae, Margarodidae, Ortheziidae, Phoenicococcidae and Pseudococcidae. Twelve genera of Hymenoptera belonging to five families were recorded as parasitoids: Aphelinidae (Aphytis, Coccophagus and Encarsia); Encyrtidae (Anagyrus, Arrhenophagus, Gyranusoidea, Habrolepis, Metaphycus and Rhopus); Eulophidae (Aprostocetus); Pteromalidae (Scutellista) and Signiphoridae (Signiphora). Four endoparasitoids were recorded from Egypt for the first time: Arrhenophagus sp. and Gyranusoidea litura Prinsloo (Encyrtidae), Aprostocetus sp. (Eulophidae) and Signiphora fax Girault (Signiphoridae).

Addison, P. & Samways, M.J. 2000. A survey of ants (Hymenoptera: Formicidae) that forage in vineyards in the Western Cape Province, South Africa. African Entomology 8(2): 251-260. [AddisoSa2000]

Notes: This study was undertaken to establish which species of ants were associated with the mealybug Planococcus ficus (Signoret) (Hemiptera: Pseudococcidae), and which species were dominant in the main vine-growing areas of the Western Cape Province, South Africa. During 1998/99, 22 vineyards were surveyed in the Stellenbosch/Paarl, Little Karoo, Worcester, Swartland, Olifants River and Hex River Valley regions, using pitfall traps to sample epigaeic ants and tuna-bait traps to sample arboreal ants. Each vineyard was sampled intensively for two consecutive weeks shortly before harvest. Forty two species of ants were recorded during the survey. The most widely distributed ant species, which were potentially dominant and associated with mealybug outbreaks in vineyards in the Western Cape Province, were Anoplolepis custodiens (F. Smith), A. steingroeveri (Forel) and Linepithema humile (Mayrn). Crematogaster peringueyi Emery, Crematogaster sp. 2 and C. melanogaster Emery are three arboreal species potentially dominant in vines only. Dominance indices for Pheidole sp. 1 and Pheidole sp. 2 were low compared to the more aggressive Anoplolepis spp. and L. humile, indicating that the former two species were not of economic significance. Edge effects occurred in five of the surveyed vineyards for three ant species. These edge effects indicate specific preferences of the ants for certain abiotic and microclimatic factors in vineyards, but could also be the result of interspecific competition.

Alencar, J.A. de 2000. [Potential pests present in the irrigated dwarf cocoa agroecosystem in the Sao Francisco valley.] Pragas potenciais presentes no agroecossistema do coqueiro anao irrigado no submedio do vale do Sao Francisco. (In Portuguese). Documentos da Embrapa Semi Arido No. 152: 12 pp. [Alenca2000]

Notes: Notes are given on potential arthropod pests of dwarf cocoa in Petrolina, Brazil. The pests considered are: Aspidiotus destructor, Hyalospila ptychis, Tetranychidae and Eriophyidae.

Allsopp, P.G. & Hogarth, D.M. 2001. I think pink ground pearls might be feeding on my crop -- what can I do? Proceedings of the 2001 Conference of the Australian Society of Sugar Cane Technologists 199-203. [AllsopHo2001]

Notes: [Conference held at held at Mackay, Queensland, Au 2001.] The morphology (at different developmental stages), development, and reproduction of pink ground pearl (Eumargarodes laingi) are reviewed. Use of the proper insecticide or cultural control mechanisms (varying length of crop cycle, fallowing, cultivation and alternative crops), resistant sugarcane cultivars; and observation of proper farm hygiene to reduce/eradicate this pest are presented. Combinations of the aforementioned practices for integrated pest management, and different strategies for various soil types are described.

Allsopp, P.G. (Ed.) & Suasa Ard, W. 2000. In: , Sugarcane Pest Management in the New Millennium. 4th Sugarcane Entomology Workshop International Society of Sugar Cane. International Society of Sugar Cane Technologists, Indooroopilly, Australia. 104 pp. [AllsopSu2000]

Notes: Thirteen papers were presented at this Workshop, under the headings: biological control of sugarcane borers; chemical and host-plant resistance for control of borers; integrated control of borers; biotechnology; biological control of sucking insects; control of whitegrubs; economic injury levels; and pest faunas and incursion management. Scale species discussed include Aulacaspis tegalensis, Ceroplastes destructor, Planococcus citri, Pseudococcus maritimus and Saccharicoccus sacchari.

Allsopp, P.G., FitzGibbon, F. & DeBarro, P.J. 2000. Development of a Pest Risk Analysis and a Pest Incursion Management Plan for the Australian Sugar Industry. 96-103 In: Allsopp, P.G. (Ed.) & Suasa Ard, W., Sugarcane Pest Management in the New Millennium. 4th Sugarcane Entomology Workshop International Society of Sugar Cane International Society of Sugar Cane Technologists, Indooroopilly, Australia. 104 pp. [AllsopFiDe2000]

Notes: A Pest Risk Analysis (PRA) undertaken for the Australian sugar industry identified 1286 species of insects and mites associated with sugarcane throughout the world. In areas to the north of Australia (Papua New Guinea, Indonesia, Malaysia, Thailand and Philippines) more than 400 potential pest species were identified, including two scale insects. We also developed a Pest Incursion Management Plan (PIMP) that complements the PRA database, and covers both incursions into commercial cropping areas and into back-yard plots of sugarcane.

Alvarez, R.C. 2001. Program for rubber. Greenfields (Philippines) p. 23-27. [Alvare2001]

Notes: The occurrence of Pseudococcidae and pest control is among the topics discussed here.

Amin, A.H., Risk Madiha, A. & Sakr, H.E.A. 2001 (1999). Factors responsible for the extinction of Chrysomphalus aonidum (L.) from citrus orchards in Egypt. Entomologica 33(1999): 441. [AminRiSa2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. Ecological studies have shown that the black armoured scale, Chrysomphalus aonidum (L.), has four overlapping annual generations on navel orange trees under conditions in Qalyubyia Governorate, Egypt. Five species of natural enemies were recorded on C. aonidum during the present work. The ectoparasitoid, Aphytis holoxanthus DeBach, was recorded for the first time from Egypt, and the other bioagents were three endoparasitoids: Aspidiotiphagus citrinus Craw., A. lounsburyi Berl. & Paoli and Habrolepis pascuorum Mercet and an entomogenous fungus, Cladosporium cladosporides (Fresen). Recently, C. aonidum appears to have become extinct in citrus orchards in many localities in Egypt. It is considered that this is due to a combination of spray timing and the effects of these five bioagents. Since 1967, the Ministry of Agriculture in Egypt has recommended the use of summer sprays against this pest, instead of autumn and winter applications. Summer sprays are less harmful to these natural enemies, which are mainly effective against the winter and spring populations of C. aonidum.

Arai, T. 2001. Effects of simulated rainfall on the survival of Pseudococcus cryptus Hempel (Homoptera: Pseudococcidae). Bulletin of the National Institute of Fruit Tree Science (35): 109-114. [Arai2001]

Notes: Potted citrus trees infested with Pseudococcus cryptus were exposed to simulated rainfall to clarify its effect on the survival of this pest species. The rainfall did not affect the survival rates of the insects. The addition of wind at a rate of 2m/s to the rainfall enhanced the mortality of the first instar nymphs. The intensity and duration of rainfall with wind affected the survival of the first instar nymphs. However, the survival of the second instar nymphs did not change by those treatments.

Aung, L.H., Leesch, J.G., Jenner, J.F. & Grafton Cardwell, E.E. 2001. Effects of carbonyl sulfide, methyl iodide, and sulfuryl fluoride on fruit phytotoxicity and insect mortality. Annals of Applied Biology 139(1): 93-100. [AungLeJe2001]

Notes: Three potential chemical fumigants, carbonyl sulfide (COS), methyl iodide (MI) and sulfuryl fluoride (SF), were tested at selected dosages on lemons (cv. Eureka) against California red scale (Aonidiella aurantii), and MI and COS were tested on nectarines (cv. Summer Bright) against codling moth (Cydia pomonella). SF at _ 40 mg litre-1 gave 100% red scale mortality but resulted in commodity phytotoxicity. Lemons treated with the highest selected dose of 80 mg litre-1 COS gave only 87% kill of red scale, but failed to reach the desired probit 9 level.

Aytas, M., Yumruktepe, R. & Mart, C. 2001. [Using pheromone traps to control California Red Scale Aonidiella aurantii (Maskell) (Hom.: Diaspididae) in the Eastern Mediterranean region.] (In Turkish; Summary In English). Turkish Journal of Agriculture and Forestry 25(2): 97-110. [AytasYuMa2001]

Notes: [Original title: Dogu akdeniz bolgesi'nde turuncgillerde zararli Kirmizi kabuklubit, Aonidiella aurantii (Maskell) (Hom.: Diaspididae) mucadelesinin yonlendirilmesinde eseysel tuzaklardan yararlanma olanaklari.] In this study, the possibility of using pheromone traps against California red scale (CRS), Aonidiella aurantii, the main pest of citrus fruits, was examined. In 1995 and 1996, the male flight periods and four peak points were determined. There was a significant correlation between numbers of trapped males and leaf/fruit CRS populations. Mass trapping was not found to be a promising method for CRS control based on effectiveness and costs.

Bajwa, G.A. & Gul, H. 2000. Some observations on insect species of Paulownia species at Pakistan Forest Institute Campus, Peshawar. Pakistan Journal of Forestry 50(1-2): 71-80. [BajwaGu2000]

Notes: Exotic Paulownia, fast growing tree species of Chinese origin, were introduced in different ecological zones of Pakistan. Studies conducted by recording the observations fortnightly on nurseries and plantations at the Pakistan Forest Institute Campus, Peshawar, showed that the Paulownia spp. (P. elongata, P. fargesii, P. fortunei, P. kawakamii, P. tomentosa) were attacked by fourteen insect species, including Drosicha stebbingii (Margarodidae). D. stebbingii appeared in epidemic form on Paulownia tomentosa and P. fortunei. The pest was managed by adopting an integrated control strategy in which practices such as destruction of eggs by soil working, banding of tree trunks and application of insecticides were combined together. All components of the integrated control strategy proved effective. Among the chemicals Buldock 25EC [beta-cyfluthrin], Mepra 50EC and Endon 35EC at 0.2% were the most effective insecticides against the pest.

Balikai, R.A. & Bagali, A.N. 2000. Population density of mealybug, Maconellicoccus hirsutus (Green) on ber (Zizyphus mauritiana Lamarck) and economic losses. Agricultural Science Digest 20(1): 62-63. [BalikaBa2000]

Notes: Studies were carried out in Bijapur, India, in 1998 to assess the population density of M. hirsutus on ber and economic losses sustained. On an average there were 80.6 colonies per plant, each colony having 17.8 individuals. Similarly 15.4 egg masses covered with white waxy mealy matter were observed per plant, each egg mass containing 90.7 orange coloured oval shaped eggs. Based on the market price of infested and healthy fruits, there was a net monetary loss of Rs. 25,800/ha accounting for 33.33 per cent loss due to mealybug infestation.

Barbagallo, S. 2000. Ricordo di Vincenzo Lupo (1908-1999). (In Italian). Atti dell'Accademia Nazionale Italiana di Entomologia 48: 67-80. [Barbag2000]

Barzman, M.S. & Daane, K.M. 2001. Host-handling behaviours in parasitoids of the black scale: a case for ant-mediated evolution. Journal of Animal Ecology 70(2): 237-247. [BarzmaDa2001]

Notes: 1. We hypothesize that differences in host-handling times among three closely related encyrtid parasitoids (Metaphycus anneckei Guerrieri & Noyes, Metaphycus hageni Daane & Caltagirone and Metaphycus lounsburyi [Howard]) are influenced by the defenses of their host, black scale (Saissetia oleae [Olivier]). Two forms of host defense were examined: (i) the scale's possession of a hard integument, and (ii) the effect of ants tending S. oleae. 2. Duration of host assessment, host rejection, drilling/oviposition and the presence or absence of host feeding were measured for M. anneckei, M. hageni and M. lounsburyi. Results show that M. anneckei oviposits through the soft ventral integument while M. hageni and M. lounsburyi oviposit through the harder dorsal integument. M. anneckei was never observed to host feed while both M. hageni and M. lounsburyi use hosts for either oviposition or host feeding. The ventral drilling location and absence of non-concurrent host feeding significantly reduced the overall host-handling time of M. anneckei, relative to M. lounsburyi and M. hageni. 3. Oviposition success of M. anneckei and M. hageni in the presence of ants was compared. An observation arena was established with the numbers of Argentine ants, Linepithema humile (Mayr) and S. oleae manipulated on potted oleanders. M. hageni and M. anneckei were placed, individually, onto the observation arena and their interaction with ants and S. oleae recorded. While 55 times 3% of M. anneckei successfully oviposited, M. hageni was unable to oviposit or host feed in the presence of foraging ants. 4. Field tests were conducted using potted oleander plants, infested with S. oleae, to determine the effect of L. humile on scale density, parasitism levels and parasitoid species composition. Ant-tended plants had significantly more scale, lower scale parasitism and lower levels of host feeding (as indicated by unknown scale mortality levels). There were also significant differences in parasitoid species composition between ant-tended and ant-excluded treatments, with the relative frequency of parasitoid species with faster host-handling times more common in the ant-tended treatment.

Beardsley, J.W. 2001 (1999). Nipaecoccus nipae (Maskell) and two apparently undescribed sibling species (Hemiptera: Coccoidea: Pseudococcidae). Entomologica 33 (1999): 49-57. [Beards2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] This paper argues that there are at least two species belonging to the Nipaecoccus nipae-species complex in Hawaii, one with white wax and the other with buff wax. These two species differ in a number of features but perhaps the most important is that the encyrtid parasitoid Pseudaphycus utilis Timberlake is only known to parasitize the white-wax species. A brief history of the two species (and another in Florida) is outlined, followed by a discussion of the morphological reasons for believing them separate. The problems of nomenclature are then outlined and it is concluded that the buff-wax species is Nipaecoccus nipae (Maskell) and that N. pseudonipae (Cockerell) is a synonym. The two other species (i.e. the white-wax species from Hawaii and the species from Florida) are then described as Nipaecoccus paranipaen, n. sp. and Nipaecoccus floridensis, n.sp. respectively.

Beggs, J. 2001. The ecological consequences of social wasps (Vespula spp.) invading an ecosystem that has an abundant carbohydrate resource. Biological Conservation 99(1): 17-28. [Beggs2001]

Notes: Introduced Vespula wasps have successfully invaded beech (Nothofagus) forests in New Zealand. By collecting honeydew, an abundant carbohydrate resource, wasps can reach high numbers. Honeydew is produced by an endemic scale insect that infests about 1 million hectares of land, 15% of New Zealand's remaining native forest. At the peak of the wasp season, honeydew beech forests had an average biomass of about 3.8 kg of introduced wasps ha-1 (10,000 workers ha-1). These abundant invaders reduce the standing crop of honeydew by more than 90% for 5 months of the year and so compete with native species (such as birds and invertebrates) that also consume honeydew. The behaviour of 3 species of native birds is affected by this reduction in honeydew. However, it is unknown whether this affects the survival or reproductive success of these birds. Nevertheless, to avoid altering the birds' behaviour, wasp density should not increase above a level where wasps revisit honeydew threads more than once every 180-400 min. Additionally, the predation rate of wasps on some invertebrate prey species is so high that the probability of an individual surviving through the wasp season is virtually nil. Hence, wasps probably reduce or eradicate populations of some invertebrates. Wasp abundance needs to be reduced by 80-90% to conserve some native invertebrate species. Wasps could also affect nutrient cycling in the honeydew beech forest community by reducing the flow of carbon to microorganisms in the phyllosphere and the soil, which ultimately could affect soil solution chemistry. Current control tools are unable to reduce wasp populations over large tracts of forest. The challenge is to identify and develop new control techniques to achieve widespread control for conservation gains. The impact of introduced social wasps provides a warning of the damage exotic ants could cause if they were to invade honeydew beech forest. New Zealand needs to be vigilant to reduce the risk of an invasion by ants or other social wasps.

Bekele, T. 2001. Insecticidal screening against enset root mealybug, Paraputo spp. AgriTopia 16(2): 2-3. [Bekele2001]

Notes: Enset [Ensete sp.] plants infested with root mealybug (Paraputo sp.) were treated with different insecticides between the 1997 and 1999 seasons, in Ethiopia. Out of the chemical insecticides tested, Phostoxin tablets and Phyrinex 48% EC resulted in mean pseudostem circumference increases of 23.23 and 32.34 cm, and in mean plant height increases of 71.09 and 58.11 cm, respectively, over the control. Phyrinex 48% EC and Phostoxin tablet provided better control of root mealybug than the other insecticides. Miraculus chalk could not be recommended for enset root mealy bug control because of its phytotoxic effect.

Ben-Dov, Y. 2001. [A note on quarantine interceptions of the San Jose scale.] (In Hebrew; Summary In English). Alon Hanotea 55: 141-142. [BenDov2001]

Notes: The San Jose scale, Quadraspidiotus perniciosus (Comstock) (Coccoidea: Diaspididae) has been frequently intercepted during 1998-2000 in Israel on commercial importation of apple, pear and quince from Austria, France, Greece, Italy, Turkey and USA (Washington State). The scale does not occur in Israel.

Ben-Dov, Y. 2001a. A new species of Puto from oaks in the eastern Mediterranean (Hemiptera, Coccoidea, Pseudococcidae). (In English; Summary In French). Revue Française d'Entomologie 23(1): 105-108. [BenDov2001a]

Notes: The female of Puto israelensis, new species, is described and illustrated.

Ben-Dov, Y. 2001b. [Pulvinaria psidii Maskell a new soft scale in Israel.] (In Hebrew; Summary In English). Alon Hanotea 55: 262-263. [BenDov2001b]

Notes: The guava scale, Pulvinaria psidii, (Hemiptera: Coccidae: Coccoidea) was found in 1999 for the first time in Israel. Host plants are listed.

Ben-Dov, Y. 2001c. Taxonomy of Aspidiotus aharonii Bodenheimer, 1924 (Hem. Coccoidea, Diaspididae) with new synonymy. Entomologist's Monthly Magazine 137: 161. [BenDov2001c]

Notes: Aspidiotus aharoni Bodenheimer, 1924, originally described from Ceratonia siliqua near Benjamina, Israel, is designated as a junior synonym of Diaspidiotus pyri Lichtenstein, 1881.

Ben-Dov, Y. 2001d. A new genus and species of soft scales (Hemiptera: Coccoidea: Coccidae) from oaks in Israel. Phytoparasitica 29(5): 400-404. [BenDov2001d]

Notes: A new genus of soft scale, Kenima, is described for Kemina galilit, a new species recorded on oaks in Israel.

Ben-Dov, Y. & Marotta, S. 2001. Taxonomy and family placement of Coccus bassi Targioni Tozzetti, 1867 (Hemiptera: Coccoidea). Phytoparasitica 29(2): 169-170. [BenDovMa2001S]

Notes: Coccus bassi Targioni Tozzetti, 1867 is assigned to the Dactylopius in the family Dactylopiidae. Notes are given on its taxonomic features. This species is compared morphologically to Coccus cacti (now Dactylopius coccus).

Ben-Dov, Y. & Marotta, S. 2001a. Stabilizing the name Aspidiotus nerii Bouché, 1833 (Hem., Coccoidea, Diaspididae). Bulletin de la Société Entomologique de France 106(2): 181-191. [BenDovMa2001Sa]

Notes: Aspidiotus nerii is placed as the valid synonym for the oleander scale. Diaspis obliquum is established as a nomen oblitum and a synonym of A. nerii.

Ben-Dov, Y. & Matile-Ferrero, D. 2001. On the distribution of Phenacoccus yerushalmi Ben-Dov (Hemiptera: Coccoidea: Pseudococcidae) in the Mediterranean Basin. Phytoparasitica 29(2): 167. [BenDovMa2001D]

Notes: This species was first described from Jerusalem, Israel, recorded on Pinus halepensis. The mealybug is widely distributed in the Mediterranean basin, but has only been found sporadically.

Ben-Dov, Y., Matile-Ferrero, D. & Gafny, R. 2000. Taxonomy of Ceroplastes rubens Maskell with description of a related new species (Hemiptera: Coccoidea: Coccidae) from Reunion, including DNA polymorphism analysis. (In English; Summary In French). Annales de la Société Entomologique de France 36(4): 423-433. [BenDovMaGa2000]

Notes: A new species, Ceroplastes reunionensis, is described from Réunion, where it was found heavily infesting Mangifera indica, Persea americana and several ornamental plants. The closely related species, C. rubens Maskell, is redescribed from topotypic material collected in Queensland, Australia. These two species differ from each other in the wax test color, body color, morphological, biological and molecular characters (RAPD). Inter- and intraspecific variation of several morphological characters of both species is discussed. Ceroplastes vinsonii Signoret, 1872, is regarded as a nomen oblitum, and, therefore, becomes a new synonym of Ceroplastes floridensis Comstock, 1881.

Ben-Yehuda, S., Hadar, D., Wysoky, M., Izhar, Y. & Swirski, E. 2001 (1999). The importance of oils in the integrated management of the pyriform sale in avocado orchards in Israel. Entomophaga 33 (1999): 273. [BenYehHaWy2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. The pyriform scale, Protopulvinaria pyriformis (Cockerell) (Hemiptera: Coccoidea: Coccidae) was discovered in Israel in 1980 and within a few years had spread to most of the avocado growing regions of the country. Several major avocado varieties (mainly Nabal) are highly susceptible to the pyriform scale. Other less susceptible avocado varieties, in the vicinity of the infected sites, can become damaged as well. The damage is caused by the secretion of large amounts of honeydew, resulting in a thick cover of sooty-mould on the leaves and on the fruit. Despite the large numbers of local natural enemies which accumulate in infested areas, as well as the establishment of the introduced encyrtid Metaphycus stanleyi Compere, which between them often cause high mortality of the scale, injury to susceptible avocado varieties has not been prevented. As a routine, application of synthetic insecticides in avocado orchards in Israel is avoided. Hence, to lower the pyriform scale populations with minimum negative consequences to the natural enemies, we studied the effect of several oils and a soap application on the pyriform scale and the activity of the natural enemies. Several mineral oils ("mayonnaise-type" 1.75% (80% A.I.) and "stylet-oil" 1% (80% A.I.)) effectively controlled the scale nymphal stages. Therefore, oil application is conducted in the spring (February-April) and in the second half of the summer (July-October), when the population consists of mainly immature stages. Cotton oil and soap (potassium salt of fatty acids) were less effective. Application of the above mentioned compounds did not interfere with the activity of the natural enemies of the scale.

Bene, G. del, Gargani, E. & Landi, S. 2000. Evaluation of plant extracts for insect control. Journal of Agriculture and Environment for International Development 94(1): 43-61. [BeneGaLa2000]

Notes: This paper reports the results of trials on the insecticidal effect of the vegetal extracts Quassia, Ryania, Derris, Urtica, Artemisia and neem (Azadirachta indica). The insects targeted included Ceroplastes rusci and Aonidia lauri. Also studied were the side effects of the various products on the common parasitoid Encarsia formosa.

Bentley, W., Rice, R.E., Brazzle, J. & Day, K. 2000. Pests of nectarine. UC Pest Management Guidelines [BentleRiBr2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.nectarine.html] This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of nectarine in the U.S., including Quadraspidiotus perniciosus.

Bentley, W.J., Coates, W.W., Hasey, J., Hendricks, C.L., Olson, W.H., Pickel, C., Sibbett, G.S. & Van Steenwyk, R.A. 2000. Pests of walnut. UC Pest Management Guidelines [BentleCoHa2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.walnuts.html] This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of walnut in the U.S., including Parthenolecanium corni, P. pruinosum, Quadraspidiotus perniciosus and Q. juglansregiae.

Bentley, W.J., Hendricks, L., Duncan, R., Silvers, C., Martin, L., Gibbs, M. & Stevenson, M. 2001. BIOS and conventional almond orchard management compared. California Agriculture 55(5): 12-19. [BentleHeDu2001]

Notes: One of the objectives of this study was to evaluate the abundance of San José scale and its associated parasitoids in Biologically Integrated Orchard Systems (BIOS) and conventional almond orchards.

Bentley, W.J., Rice, R.E. & Day, K.R. 2000 (1998). Pests of plum. UC Pest Management Guidelines [BentleRiDa2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.plum.html] This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of plum in the U.S., including Parthenolecanium corni, Epidiaspis leperii and Quadraspidiotus perniciosus.

Bentley, W.J., Rice, R.E., Beede, R.H. & Daane, K. 2000. Pests of pistachio. UC Pest Management Guidelines. [BentleRiBe2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.pistachios.html] This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of pistachio in the U.S., including Coccus hesperidum, Saissetia oleae, Parthenolecanium corni and P. pruinosum.

Bentley, W.J., Zalom, F., Granett, J., Smith, R.J., Varela, L. & Purcell, A. 2000. Pest of grape. UC Pest Management Guidelines [BentleZaGr2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.grapes.html] This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of grape in the U.S., including Pseudococcus maritimus, P. viburni and P. longispinus.

Bergelson, J., Dwyer, G. & Emerson, J.J. 2001. Models and data on plant-enemy coevolution. Annual Review of Genetics 35: 469-499. [BergelDwEm2001]

Notes: Although coevolution is complicated, in that the interacting species evolve in response to each other, such evolutionary dynamics are amenable to mathematical modeling. In this article, we briefly review models and data on coevolution between plants and the pathogens and herbivores that attack them. We focus on "arms races," in which trait values in the plant and its enemies escalate to more and more extreme values. Untested key assumptions in many of the models are the relationships between costs and benefits of resistance in the plant and the level of resistance, as well as how costs of virulence or detoxification ability in the enemy change with levels of these traits. A preliminary assessment of these assumptions finds only mixed support for the models. What is needed are models that are more closely tailored to particular plant-enemy interactions, as well as experiments that are expressly designed to test existing models. Black pineleaf scale, beech scale, armored scale insect and pinyon needle scale are mentioned as examples of empirical studies of local adaptation.

Bernal, J.S., Luck, R.F., Morse, J.G. & Drury, M.S. 2001. Seasonal and scale size relationships between citricola scale (Homoptera: Coccidae) and its parasitoid complex (Hymenoptera: Chalcidoidea) on San Joaquin Valley citrus. Biological Control 20(3): 210-221. [BernalLuMo2001]

Notes: The phenology of citricola scale, Coccus pseudomagnoliarum (Kuwana), and its associated parasitoid complex were studied on citrus in the San Joaquin Valley of central California over the period April 1995-March 1997. A total of 10,237 parasitoid specimens of 10 species were collected. Two of these species, Marietta mexicana (Howard) and Encyrtus lecaniorum (Mayr), each recovered from individually isolated scales, represent new parasitoid records for citricola scale. A third species, Encarsia citrinus citrinus (Craw), may represent a new parasitoid record, but this requires further confirmation because a single (male) specimen was recovered from individually isolated scales. The three most dominant parasitoid species, Coccophagus lycimnia (Walker), Metaphycus helvolus (Compere), and Metaphycus Luteolus (Timberlake), accounted for the majority (>97%) of the specimens recovered. In contrast to the situation on citrus in southern California, where citricola scale is under effective biological control and is very rarely seen, citricola scale on citrus in the San Joaquin Valley is reemerging as a major pest, especially in groves employing integrated pest management with minimal use of broad-spectrum insecticides. Possible reasons uncovered in this study for the lack of effective biological control of citricola scale in the San Joaquin Valley include: (i) reduced presence of Metaphycus spp. because of hyperparasitism by the heteronymous hyperparasitoid C. lycimnia; (ii) absence of alternate hosts for those species of Metaphycus present; and (iii) absence of hosts of suitable size for Metaphycus at critical times of the year. Recommendations for improving the level of biological control in the San Joaquin Valley are discussed.

Bertschy, C., Turlings, T.C.J., Bellotti, A.C. & Dorn, S. 2001. The role of mealybug-induced cassava plant volatiles in the attraction of the encyrtid parasitoids Aenasius vexans and Apoanagyrus diversicornis. Journal of Insect Behavior 14(3): 363-371. [BertscTuBe2001]

Bilog Obra, G.P. & Morallo Rejesus, B. 2000. Biological studies of Aphytis sp. nr. chrysomphali (Hymenoptera: Aphelinidae). Philippine Entomologist 14(2): 137-147. [BilogOMo2000]

Notes: The parasitoid Aphytis sp. nr. chrysomphali was collected from coconut scale, California red scale (Aonidiella aurantii) and Florida red scale (Chrysomphalus ficus) attacking citrus in August 1989 from Villaverde, Nueva Vizcaya, Philippines. The biology of the parasitoid on coconut scale, Aspidiotus destructor Signoret, reared on squash was studied. The total developmental period of the uniparental (thelytokous) parasitoid from egg deposition to adult emergence ranges from 13-15 days with a mean of 14.18+0.90.

Biswas, J. & Ghosh, A.B. 2000. Biology of the mealybug, Planococcus minor (Maskell) on various host plants. Environment & Ecology 18(4): 929-932. [BiswasGh2000]

Notes: The biology of the mealybug, Planococcus minor (Maskell) was studied in the laboratory on different host plants. The average duration of male and female nymphs respectively were 21.42 and 16.17 days on Ixora signaporensis, 16.73 and 17.49 days on Glycine max and 21.30 and 21.80 days on Acalypha wilkesiana. The preoviposition and oviposition periods and fecundity were 8.25 and 9.25 days and 65.75 eggs on I. signaporensis, 6.25 and 8.25 days and 132.50 eggs on G. max and 7.25 and 9.25 days and 138.75 eggs on A. wilkesiana.

Blachinski, D. & Dreeshpon, Y. 2000. [Spring control of the citrus mealybug (Planococcus citri).] (In Hebrew). Alon Hanotea 54(8): 316-318. [BlachiDr2000]

Blumberg, D. & Van Driesche, R.G. 2001. Encapsulation rates of three encyrtid parasitoids by three mealybug species (Homoptera: Pseudococcidae) found commonly as pests in commercial greenhouses. Biological Control 22: 191-199. [BlumbeVa2001]

Notes: Encapsulation rates of the parasitoids Leptomastix dactylopii Howard, Leptomastix epona (Walker), and Anagyrus fusciventris (Girault) (Homoptera: Encyrtidae) by the mealybugs Planococcus citri Risso, Pseudococcus viburni (Signoret), and Pseudococcus longispinus (Targioni-Tozzetti) (Homoptera: Pseudococcidae) were studied under controlled laboratory conditions.

Blumberg, D., Ben-Dov, Y. & Mendel, Z. 2001 (1999). The citriculus mealybug, Pseudococcus cryptus Hempel, and its natural enemies in Israel: History and present situation. Entomologica 33 (1999): 233-242. [BlumbeBeMe2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The citriculus mealybug, Pseudococcus cryptus Hempel, was first discovered in Israel in 1937 and very rapidly became a key pest of citrus. However, since the early 1940s, the mealybug population has sharply decreased. This occurred in parallel with the establishment of the introduced parasitoid Clausenia purpurea Ishii, which was then believed to be the main cause of the biological control of the mealybug. Since the late 1980s, outbreaks of P. cryptus have been recorded mainly in new citrus varieties, such as red grapefruits, pomelo, "sweety" and several peeling varieties. The current outbreaks are probably related to the susceptibility of these mentioned varieties to P. cryptus, and to the adverse effects of Insect Growth Regulators to coccinellid predators, especially Scymnus spp. The introduced C. purpurea and two other local encyrtid parasitoids, Leptomastix near algirica and Anagyrus diversicornis Mercet, rarely emerged from samples of P. cryptus collected during 1996-1998. Four further parasitoid species were introduced into Israel during 1996-1997 against P. cryptus: from central Asia, the platygasterids Allotropa burrelli Muesebeck and A. convexifrons Muesebeck and the encyrtid, Pseudaphycus malinus Gahan; and from Japan, Anagyrus sawadai Ishii. A. convexifrons and A. sawadai successfully parasitized P. cryptus and, therefore, were released in the field but only A. sawadai has so far been recovered. A considerable reduction in population densities of the pest has been recorded since May, 1998, in the major release site of the latter species.

Bogo, A. & Mantle, P. 2000. Oligosaccharides in the honeydew of Coccoidea scale Insects: Coccus hesperidum L. and a new Stigmacoccus sp. in Brazil. Anais da Sociedade Entomologica do Brasil 29(3): 589-595. [BogoMa2000]

Notes: Analysis of the honeydew from an as yet undescribed, though distinctive, Brazilian Stigmacoccus sp. (near S. asper Hempel) by paper chromatography, Fast atom bombardment (FAB-MS) and Gas chromatography-mass spectrometry (GC-MS) identified fructose and glucose as monosaccharides and sucrose, maltose, trehalulose, trehalose and a hexose-hexitol as disaccharides. Erlose and glucosyl erlose have been identified as the tri- and tetra-saccharides in Stigmacoccus sp. and characterised for the first time in scale insects by modern techniques of linkage analysis. The same erlose oligosaccharides were recognised in honeydew of the common scale insect Coccus hesperidum L., together with the pentamer of this series, maltosyl erlose, therefore recognising that specific metabolic transformations of sugars into this oligomeric series occur rather widely in scale insects.

Bogo, A., Watson, G.W., Mantle, P.G. & Mottana G.M. 2001 (1999). Honeydew sugars eliminated by Stigmacoccus sp. nr. asper Hempel (Hemiptera: Margarodidae) feeding on leguminous trees in Brazil. Entomologica 33(1999): 275-278. [BogoWaMa2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The sooty mould coating the trunks of mature trees of Schizolobium excelsum in Brazil was found to be associated with honeydew being eliminated by an undescribed species of margarodid near Stigmacoccus asper Hempel. Analysis of the honeydew sugars by paper chromatography revealed a complex composition. The principal sugar was sucrose, but there were significant amounts of fructose, glucose and three components identified as di-, tri- and tetrasaccharides. The disaccharides were maltose, trehalose, trehalulose and a hexose-hexitol. The other, apparently novel, pair of oligosaccharides were composed of glucose(s) 1,4 linked to the glucose of sucrose. The sugar composition of the tree sap was also determined and found to be glucose and sucrose only. The findings, therefore, imply significant and novel metabolic transformations of sugars by the scale insect and/or its microbial symbionts.

Bogran, C.E., Heinz, K.M. & Ciomperlik, M.A. 2002. Interspecific competition among insect parasitoids: Field experiments with whiteflies as hosts in cotton. Ecology 83(3): 653-668. [BogranHeCi2002]

Bohidar, K. 2001 (1999). The taxonomic relationship of some Coccoidea (Hemiptera) based on male characteristics. Entomologica 33(1999): 165. [Bohida2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. Male characters of species from five different families were studied and the taxonomic status of the families is described. A key of characters was presented for adult males. The quantitative evaluation method was applied to assess the relationship between the families. All the characters included were regarded as being of equal importance and those shared by any one family exhibited some differences, justifying the recognition of the taxon as a distinct family. It was found that the Margarodidae were the most primitive group followed by Pseudococcidae. The Diaspididae showed the maximum number of specialised characters, with the Coccidae and Dactylopiidae falling between the Diaspididae and the Pseudococcidae. The Coccidae and Dactylopiidae shared an equal number of characters with the Pseudococcidae and are considered to be close to each other.

Bongiorni, S., Mazzuoli, M., Masci, S. & Prantera, G. 2001. Facultative heterochromatization in parahaploid male mealybugs: involvement of a heterochromatin-associated protein. Development 128(19): 3809-3817. [BongioMaMa2001]

Notes: The behavior of chromosomes during development of the mealybug Planococcus citri provides one of the most dramatic examples of facultative heterochromatization. In male embryos, the entire haploid paternal chromosome set becomes heterochromatic at mid-cleavage. Male mealybugs are thus functionally haploid, owing to heterochromatization (parahaploidy). To understand the mechanisms underlying facultative heterochromatization in male mealybugs, we have investigated the possible involvement of an HP-1-like protein in this process. HP-1 is a conserved, nonhistone chromosomal protein with a proposed role in heterochromatinization in other species. It was first identified in Drosophila melanogaster as a protein enriched in the constitutive heterochromatin of polytene chromosome. Using a monoclonal antibody raised against the Drosophila HP-1 in immunoblot and immunocytological experiments, we provide evidence for the presence of an HP-1-like in Planococcus citri males and females. The results indicate a role for an HP-1-like protein in the facultative heterochromatization process.

Botton, M., Hickel, E.R., Soria, S.J. & Teixeira, I. 2000. [Bioecology and control of ground pearl Eurhizococcus brasiliensis (Hempel, 1922) (Hemiptera: Margarodidae) in vine culture.] Bioecologia e controle da Pérola-da-terra Eurhizococcus brasiliensis (Hemiptera: Margarodidae) na cultura da videi (In Portuguese). Circular Técnica (EMBRAPA: Empresa Brasileira de Pesquisa Agropecuária) 27: 23 pp. [BottonHiSo2000]

Notes: E. brasiliensis is briefly described and photos are provided. A list of about 70 of its host plants, a description of the damage it inflicts on its hosts, its geographical distribution, and biological, cultural and chemical control are discussed.

Bribosia, E., Champagne, R. & Bylemans, D. 2000. [Useful organisms in orchards: the lady birds.} Nuttige organismen in boomgaarden: de lieveheersbeestjes. (In Dutch). Fruitteelt Nieuws 13(22) 26-28. [BribosChBy2000]

Notes: The most important Coccinellidae occurring in orchards are described, including Adalia bipunctata, Synharmonia conglobata [Oenoplia conglobata], and A. decempunctata (predators of aphids), Chilocorus bipustulatus and Exochomus quadripustulatus (predators of scales), and Stethorus punctillum (predator of mites). The effects of ecological factors (feed preference, habitat - especially vegetation layer, and natural enemies) on behaviour are also described.

Broschat, T.K. & Weissling, T.J. 2001. Susceptibility of lantana cultivars to Orthezia insignis. HortTechnology 11(3): 460-462. [BroschWe2001]

Notes: The greenhouse orthezia (Orthezia insignis) is a serious and widespread pest of cultivated lantanas (Lantana sp.) in warmer regions of the world. Forty species and cultivars of lantanas were screened for their relative susceptibility to this insect pest. Results showed that two Florida native lantanas, pineland lantana (L. depressa) and buttonsage (L. involucrata), were highly susceptible to infestation, with trailing lantana (L. montevidensis) and its cultivars and hybrids being somewhat less susceptible. Shrub lantana (L. camara) and its cultivars and hybrids were the least susceptible to greenhouse orthezia infestation, but some of these varieties are rather unattractive as landscape ornamentals and can become serious weeds.

Böhmer, H. 2000. Insect dyes. HALI: Carpet, Textile and Islamic Art 113: 57-60. [Bohmer2000]

Notes: Scale species mentioned include Dactylopius coccus, Kerria lacca, Porphyrophora hameli, P. polonica and P. tritici.

CAB International. 2000. Phenacoccus madeirensis Green. Distribution Maps of Pests Map no. 607: 4 pp. [CABI2000]

Notes: Published in the U.K. by the International Institute of Entomology, an agency of CAB International; map showing distribution of this sp. worldwide; countries listed with references to records; attacks cassava (Manihot esculenta) and many other crops including aubergine (Solanum melongena), oat (Avena sativa), cotton (Gossypium spp.), Coleus, Hibiscus, potato (Solanum tuberosum), Capsicum annuum and Citrus spp.

CAB International. 2000a. Sphaeraspis vitis (Philippi). Distribution Maps of Pests Map no. 608: 2 pp. [CABI2000a]

Calatayud, P.A., Auger, J., Thibout, E., Rousset, S., Caicedo, A.M., Calatayud, S., Buschmann, H., Guillaud, J., Mandon, N. & Bellotti, A.C. 2001. Identification and synthesis of a kairomone mediating host location by two parasitoid species of the cassava mealybug Phenacoccus herreni. Journal of Chemical Ecology 27(11): 2203-2217. [CalataAuTh2001]

Notes: Two encyrtid species, Acerophagus coccois and Aenasius vexans, parasitoids of the cassava mealybug Phenacoccus herreni use a contact kairomone from the body surface of their host as a host-location stimulant. The kairomone was synthesized and identified as O-caffeoylserine based on a combination of chromatographic methods. The synthetic compound was determined to be active.

Calatayud, P.A., Seligmann, C.D., Polania, M.A. & Bellotti, A.C. 2001. Influence of parasitism by encyrtid parasitoids on the feeding behaviour of the cassava mealybug Phenacoccus herreni. Entomologia Experimentalis et Applicata 98(3): 271-278. [CalataSePo2001]

Notes: Three encyrtid parasitoids Apoanagyrus (Epidinocarsis) diversicornis, Aenasius vexans, and Acerophagus coccois (Hymenoptera: Encyrtidae) are used to control the cassava mealybug Phenacoccus herreni Cox & Williams (Sternorrhyncha: Pseudococcidae), an important pest of cassava in South America. The influence of parasitism on the feeding behaviour of mealybugs was studied by observing honeydew secretion and by the electrical penetration graph technique (EPG, DC-system). Honeydew secretions were observed after parasitism until mummy transformation. No strong EPG parameter differences were found between parasitised and control insects. All results indicated that parasitised mealybugs keep feeding on the phloem sap after parasitism until mummy transformation. The main influence of parasitism on EPG parameters is the appearance of a new pattern resembling the E2 pattern at the extracellular level and labelled H. This pattern was also produced with control insects located on an unfavourable feeding site and could be associated with a stress response. It might be related to the still unclear E(c) pattern of aphids. The relationship of H to stylet activities is discussed.

Cantini, R. & Battisti, A. 2001. Impact and control of the cone tortricid Pseudococcux tessulatana (Staudinger), damaging the cone crop of a selected clone of cypress (Cupressus sempervirens L.) in Italy. Anzeiger für Schädlingskunde 74(4): 107-110. [CantinBa2001]

Notes: The green cypress (Cupressus sempervirens) is of great interest for ornamental, reforestation and windbreak use in the whole Mediterranean basin. In Italy, seed material selected for resistance to the fungus Seiridium cardinale is produced in seed orchards by controlled crosses of parent trees. The insect pest showing the highest impact on seed cone production is Pseudococcux tessulatana (Lep.; Tortricidae), which attacks cones during the initial growth period as well as full-grown cones. The impact on the seed cone crop of the tortricid was estimated on a clone patented for its resistance to cypress canker (Agrimed 1). The attack was inversely related to the cone crop, as it concerned 36.7% of cones in 1999 (high crop year) and the 66% in 2000 (low crop year). In both years, about 90% of the surveyed branches revealed cones attacked by P. tessulatana by the 1st life-cycle larvae, whereas only 40% of branches were also attacked by the larvae of the 2nd life cycle. The highest attack rate per branch was always observed on branches bearing a low number of cones. The potential of two control methods against P. tessulatana to protect cones which result from crossing a mother tree "Agrimed 1" with selected father trees was also evaluated in 2000. The protection given by a sleeve surrounding the branch was almost complete (0.4% cone mortality), whereas a cone mortality of 24.3% was observed on branches treated by the insecticide diflubenzuron. Sleeves appear to be useful to protect branches on which special crosses were done, but are expensive and time-consuming and may favour the attack of the mealybug Planococcus vovae inside the sleeve.

Caprile, J., Varela, L., Pickel, C., Coates, W.W., Bentley, W.J. & Vossen, P.M. 2000. Pests of apple. UC Pest Management Guidelines [CaprilVaPi2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.apples.html] This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of apple in the U.S., including Epidiaspis leperii and Quadraspidiotus perniciosus.

Chan, Y.K., Subhadrabandhu, S. (Ed.) & Chairidchai, P. 2000. Status of the pineapple industry and research and development in Malaysia. Acta Horticulturae No. 529: 77-83. [ChanSuCh2000]

Notes: [Proceedings of the Third International Pineapple Symposium, Pattaya, Thailand, 17-20 November 1998.] Malaysia, once ranked as one of the top three pineapple producers in the world in the 1960s and early 1970s, has only a relatively modest industry today. However, the fresh pineapple export had picked up recently to 40000 tonnes worth RM 10 million in 1997. With the introduction of the new hybrid Josapine in 1996, the market for fresh pineapple is expected to increase further. Current research includes a focus on the management of mealybug closterovirus red wilt.

Charles, J.G., Froud, K.J. & Henderson, R.C. 2001 (1999). Morphological variation in the mealybugs Pseudococcus calceolariae and P. similans (Hemiptera: Coccoidea: Pseudococcidae). Entomologica 33(1999): 165. [CharleFrHe2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. Examination of slide-mounted laboratory-reared daughters from isolines of Pseudococcus calceolariae (Maskell) and P. similans (Lidgett) mothers collected from two widely separated locations in the North Island of New Zealand has thrown doubt on the validity of the defining characters of the species. For P. similans, the number and position of oral rim tubular ducts varied widely and sometimes fell outside the defined species limits. In addition, the characteristic `stout abdominal setae' were lost in the F1 generation. The combinations of leg-length and number of oral rim tubular ducts which separate P. calceolariae (Maskell) from P. similans co-existed in cohorts of F1 sisters and both character-sets could be manipulated by changing the temperature at which the mealybugs developed. Mating studies are proposed to investigate whether the two species are valid, or whether they merely represent the phenotypic extremes of one widely polymorphic species.

Chatterjee, H., Ghosh, J. & Senapati, S.K. 2000. Influence of important weather parameters on population fluctuation on major insect pest of mandarin orange (Citrus reticulata Blanco) at Darjeeling district of West Bengal (India). Journal of Entomological Research. New Delhi 24(3): 229-233. [ChatteGhSe2000]

Notes: Saissetia coffeae shows a positive association with maximum temperature, R.H. and rainfall, but an inverse relation with mimimum temperature. An identical relationship was observed with Coccus hesperidum for maximum and minimum temperatures.

Chen, Y.P. & Li, W.L. 2001. [Study on the major host plant species of lac insect, wax scale and gall aphids.] (In Chinese). Forest Research 14(1): 100-105. [ChenLi2001]

Notes: Lac insect, wax scale and gall aphids are the important resource insects in China. The host plant is not only the habitant and reproductive sites for resource insects but also their food resource. The species of host plant are various, and some of them are excellent hosts. The host plant species were extensively collected from the major production regions of lac, wax and Chinese gallnut, and then reproduced in Jingdong of Yunnan which is located in the southern subtropical region. More than 200 species, which belong to 98 genera and 36 families of host plants of lac insect, wax scale and gall aphids have been collected over many years. These species are valuable for studying these resource insects and their host plants.

Chirinos-Torres, L., Geraud-Pouey, F., Chirinos, D.T., Fernandez, C., Guerrero, N., Polanco, M.J., Fernandez, G. & Fuenmayor, R. 2000. [Effect of insecticides on Capulinia sp. near jaboticabae von Ihering (Hemiptera: Eriococcidae) and its natural enemies in Mara county, Zulia State, Venezuela.] (In Spanish; Summary In English). Boletín de entomología venezolano 15(1): 1-16. [ChirinGeCh2000]

Notes: [Original title: Efecto de insecticidas sobre Capulinia sp. cercana a jaboticabae von Ihering (Hemiptera: Eriococcidae) y sus enemigos naturales en el municipio Mara, estado Zulia, Venezuela.] The effect of several insecticide treatments on the guava cottony scale, Capulinia sp., and its natural enemies, was assessed under field conditions in Mara county, Zulia state, during April-September 1995.

Claps, L.E. 2000. [Redescription of five species of Diaspididae (Hemiptera, Coccoidea) of the Neotropical region.] Redescripción de cinco especies de Diaspididae (Hemiptera, Coccoidea) de la región neotropical. (In Spanish; Summary In English). Revista Brasileira de Entomologia 44(3/4): 91-95. [Claps2000]

Notes: Acutaspis paulista, Dinaspis ichesii, D. lahillei, Protargionia larreae and Targionia fabianae are described and illustrated. Synonymy, plants hosts and biology (when known) are given.

Claps, L.E. & Terán, A.L. 2001. [Diaspididae (Hemiptera: Coccoidea) associated with citrus in the province of Tucuman (Argentina). (In Spanish; Summary In English). Neotropical Entomology 30(3): 391-402. [ClapsTe2001]

Notes: [Original title: Diaspididae (Hemiptera: Coccoidea) asociadas a cítricos en la provincia de Tucumán (República Argentina).] Aonidiella aurantii, Aspidiotus nerii, Borchseniaspis palmae, Cornuaspis beckii, Chrysomphalus aonidum, C. dictyospermi, Insulaspis gloverii, Mycetaspis personata, Parlatoria cinerea, Parlatoria pergandii, Pinnaspis aspidistrae and Unaspis citri are discussed on citrus plants in Tucumán, Argentina. A key for the recognition of the species, as well as the synonymy, characterization and illustration are also included. Data on economic importance and host plants are provided.

Claps, L.E., Wolff, V.R.S. & González, R.H. 2001 (1999). Catálogo de las especies de Diaspididae (Hemiptera: Coccoidea) nativas de Argentina, Brasil y Chile. (In Spanish; Summary In English). Insecta Mundi 13(3/4): 239-256. [ClapsWoGo2001]

Notes: [Publication distributed and postmarked 2001; date printed on issue is 1999.] 96 species of armoured scale insects (Diaspididae) described from Brazil, Argentina and Chile are reviewed in terms of synonymy, location of plant material, host plants, distribution within these southern hemisphere countries and bibliographical references.

Claps, L.E., Wolff, V.R.S. & González, R.H. 2001a. [Catalog of the exotic species of Diaspididae (Hemiptera: Coccoidea) from Argentina, Brazil and Chile.] (In Spanish; Summary In English). Revista de la Sociedad Entomológica Argentina 60(1/4): 9-34. [ClapsWoGo2001a]

Notes: [Original title: Catálogo de las Diaspididae (Hemiptera: Coccoidea) exóticas de la Argentina, Brasil y Chile.] 87 exotic species of armored scale insects from Brazil (69), Argentina (62) and Chile (29) are listed. Each species is reviewed in terms of host plants, geographical distribution, bibliographical references and economic importance. Aonidiella, Quadraspidiotus and Parlatoria are mentioned as important for quarantine.

Colen, K.G.F., Santa Cecilia, L.C.V., Moraes, J.C. & Reis, P.R. 2000. [Effects of different temperatures upon the biology of the mealybug Dysmicoccus brevipes (Cockerell, 1893) (Hemiptera: Pseudococcidae).] (In Portuguese; Summary In English). Revista Brasileira de Fruticultura 22(2): 248-252. [ColenSaMo2000]

Notes: [Original title: Efeitos de diferentes temperaturas sobre a biologia da cochonilha pulverulenta Dysmicoccus brevipes (Cockerell, 1893) (Hemiptera: Pseudococcidae).] The biology of the mealybug Dysmicoccus brevipes (Cockerell, 1893) was studied at four temperatures: 20, 25, 30 and 35̊C, aiming at the determination of its thermal requirements as basic information for pineapple pest management programs. The experiment was carried out at the Entomology Laboratory of the Centro Tecnologico do Sul de Minas (Technological centre of the south of Minas) -- CTSM/EPAMIG in Lavras-MG, during the period of June, 1996 to July, 1998, in a climate chamber. The insects were maintained on leaf fragments of pineapple cv. Smooth Cayenne. The length of the nymphal stages was inversely proportional to the temperature. Nymphal development was not completed at 35̊C. The highest longevities were found at the temperature of 20̊C for females and at 20 and 25̊C for males. The inferior thermal threshold for development (base temperature) was 12.1; 13.5; 12.8 and 12.8̊C for the 1st and 2nd instar nymphs, cocoon and nymphal periods, respectively. The thermal constants (K) for the nymphal phase of males was 393.4 degree-days and for females 605.6 degree-days. With the exception of the first instar, all the thermal constants obtained for females were superior to those determined for males.

Cook, L. 2001a (1999). Apiomorpha: an extraordinary scale insect genus. Entomologica 33: 166. [Cook2001a]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. Apiomorpha is a gall-inducing eriococcid genus which is endemic to Australia, where it is restricted to Eucalyptus. Females of most species induce large and spectacular galls which generally have species-specific morphologies. Apiomorpha is one of the most chromosomally diverse of all animal genera. Closely related species usually have very different karyotypes and there is extensive chromosomal variation between populations within most morphologically defined species. Some, but not all, of the karyotypic variation within morphospecies probably represents species-level differences. A phylogeny of Apiomorpha based on DNA sequence data has been proposed, onto which chromosomal and morphological traits have been mapped. Both fusion and fragmentation appear to be playing a role in chromosomal evolution in this genus.

Cook, L. & Gullan, P.G. 2001 (1999). Are the enlarged ducts of Eriococcus (Hemiptera: Coccoidea: Eriococcidae) plesiomorphic? Entomologica 33(1999): 59-66. [CookGu2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Borchsenius (1948) separated the genus Eriococcus Targioni-Tozzetti (Eriococcidae) from Acanthococcus Signoret and Gossyparia Signoret on the basis of the occurrence of enlarged ducts in the adult female of the type-species, E. buxi (Fonscolombe). Enlarged ducts are found also in another Palaearctic species, a Chilean eriococcid and several Australian species of Eriococcus. The enlarged ducts of Eriococcus are similar in appearance and distribution to the large oral rim ducts of Ferrisia Fullaway (Pseudococcidae) and the dorsal tubercle ducts of Ceronema Maskell and some species of Pulvinaria Targioni-Tozzetti (Coccidae). It is argued that the enlarged ducts in taxa from each of these three families are homologous and therefore may be plesiomorphic for the Eriococcidae. If so, the possession of enlarged ducts is not of itself sufficient to justify the separation of Eriococcus as defined by Borchsenius.

Cook, L., Gullan, P. & Trueman, H. 2001 (1999). A putative molecular phylogeny of nine scale insect "families" and its implications. Entomologica 33(1999): 166. [CookGuTr2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. An incomplete and only partially tested molecular phylogeny of the scale insects based on 18S rDNA using sample taxa from the Margarodidae, Ortheziidae, Pseudococcidae, Coccidae, Kerriidae, Eriococcidae, Diaspididae, Dactylopiidae and Asterolecaniidae is presented. Aphids and psyllids have been included as outgroup taxa. The general topology is discussed in relation to the evolution of the scale insects. Several well-supported clades have major implications for the classification and taxonomy of the Eriococcidae. A phylogeny of the eriococcids based on sequence data from the mitochondrial gene cytochrome oxidase subunit II is in general agreement with the nuclear gene-based tree.

Cook, L.G. 2000. Extraordinary and extensive karyotypic variation: A 48-fold range in chromosome number in the gall-inducing scale insect Apiomorpha (Hemiptera: Coccoidea : Eriococcidae). Genome 43(2): 255-263. [Cook2000]

Notes: Chromosome number reflects strong constraints on karyotype evolution, unescaped by the majority of animal taxa. Although there is commonly chromosomal polymorphism among closely related taxa, very large differences in chromosome number are rare. This study reports one of the most extensive chromosomal ranges yet reported for an animal genus. Apiomorpha Rubsaamen (Hemiptera: Coccoidea: Eriococcidae), an endemic Australian gall-inducing scale insect genus, exhibits an extraordinary 48-fold variation in chromosome number with diploid numbers ranging from 4 to about 192. Diploid complements of all other eriococcids examined to date range only from 6 to 28. Closely related species of Apiomorpha usually have very different karyotypes, to the extent that the variation within some species-groups is as great as that across the entire genus. There is extensive chromosomal variation among populations within 17 of the morphologically defined species of Apiomorpha indicating the existence of cryptic species-complexes. The extent and pattern of karyotypic variation suggests rapid chromosomal evolution via fissions and (or) fusions. It is hypothesized that chromosomal rearrangements in Apiomorpha species may be associated with these insects' tracking the radiation of their species host genus, Eucalyptus.

Cook, L.G. 2001. Extensive chromosomal variation associated with taxon divergence and host specificity in the gall-inducing scale insect Apiomorpha munita (Schrader) (Hemiptera: Sternorrhyncha: Coccoidea: Eriococcidae) Biological Journal of the Linnean Society 72(2): 265-278. [Cook2001]

Notes: Apiomorpha Rubsaamen (Hemiptera: Coccoidea: Eriococcidae) is one of the most chromosomally diverse of all animal genera. There is extensive karyotypic variation within many of the morphologically defined species, including A. munita (Schrader) which is here reported to have diploid chromosome counts ranging from 6 to more than 100. Each of the three morphologically defined subspecies of A. munita also displays considerable chromosomal variation: A. m. tereticornuta Gullan (2n =6, 8, 20, 22 or 24), A. m. malleensis Gullan (2n =6, 20, 22, 24 or 26), and A. m. munita (Schrader) (2n=54 or >100). Apiomorpha munita appears to occur only on eucalypts of the informal subgenus Symphyomyrtus, with each of the subspecies of A. munita restricted to discrete symphyomyrt sections. Several different karyotypic forms within each subspecies of A. munita appear to be restricted to only one or a few eucalypt species or series. The association between apparent host specificity and chromosomal rearrangements in A. munita suggests that both may be playing an active role in taxon divergence in Apiomorpha.

Cook, L.G., Gullan, P.J. & Stewart, A.C. 2000. First-instar morphology and sexual dimorphism in the gall-inducing scale insect Apiomorpha Rubsaamen (Hemiptera: Coccoidea: Eriococcidae). Journal of Natural History 34(6): 879-894. [CookGuSt2000]

Notes: Sexual dimorphism among crawlers of the scale insect family Eriococcidae is reported for the first time. The general morphology of crawlers of the gall-inducing genus Apiomorpha (Eriococcidae) is presented and sexual dimorphism described. Sexual dimorphism appears to be associated with differential dispersal and settling-site preference of the sexes during the crawler stage. First-instar males of the A. pharetrata and A. munita species-groups settle only on the galls induced by their mothers or, in the case of A. munita, also galls of nearby females, whereas female crawlers disperse. Female crawlers of all species of Apiomorpha, and male crawlers of most species, are well suited for air-borne dispersal. It is suggested that sexual dimorphism among crawlers of Apiomorpha, and some other scale insects, is the result of loss or reduction of those morphological features associated with dispersal. In addition, male crawlers of some species of Apiomorpha have sensory structures which may assist in the detection of sex-specific settling sites.

Costa, H., Cowles, R., Hartin, J., Kido, K. & Kaya, H. 2000. Pests of turfgrass. UC Pest Management Guidelines [CostaCoHa2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.turfgrass.html] This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of turfgrass in the U.S., including Odonaspis ruthae.

Coviello, R. & Bentley, W.J. 2000. Pests of fig. UC Pest Management Guidelines [CovielBe2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.figs.html] This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of fig in the U.S., including Lepidosaphes conchiformis.

Cravedi, P. 2000 (2001). Integrated peach production in Italy: objectives and criteria. (In German; Summary In French, Russian, Spanish). Pflanzenschutz Nachrichten Bayer 53(2-3): 177-197. [Craved2000]

Notes: Research has been conducted into integrated pest control for the peach tree in parts of Northern Italy since the early '70's. Research into integrated pest control of the peach tree has been conducted on the oriental fruitmoth, Cydia [Grapholita] molesta, on the green peach aphid, Myzus persicae, on scales, Pseudaulacaspis pentagona and Quadraspidiotus [Diaspidiotus] perniciosus, on thrips, in particular Frankliniella occidentalis, and on the Mediterranean fruit fly, Ceratitis capitata. The effect of insecticides on the natural beneficials of key pests were taken into account in the choice of insecticides.

Cueva, B., Izquierdo, G., Crespo, J.F. & Rodriguez, J. 2001. Unexpected spice allergy in the meat industry. Journal of Allergy and Clinical Immunology 108(1): 144. [CuevaIzCr2001]

Daane, K.M., Barzman, M.S., Caltagirone, L.E. & Hagen, K.S. 2000. Metaphycus anneckei and Metaphycus hageni: two discrete species parasitic on black scale, Saissetia oleae. BioControl 45(3):269-284. [DaaneBaCa2000]

Notes: Parasitized black scale, Saissetia oleae, were collected from olives (Olea europaea L.) in Spain and from this material an encyrtid parasitoid was reared that is taxonomically similar to Metaphycus anneckei Guerrieri and Noyes. Differences were noted between the Spanish Metaphycus nr. sp. anneckei and material described as M. anneckei that is resident to California. Although dissimilar in few morphological characters, the ovipositor length and shape of the male genitalia and the antennal club of both sexes can be used to discriminate between M. anneckei resident to California and the Spanish material, which was subsequently named Metaphycus hageni Daane and Caltagirone. To determine if morphological differences correspond to biological differences, reproductive and oviposition behaviours of Californian M. anneckei and Spanish M. hageni were studied. Mating was not observed between individuals of the two types and only male progeny were produced in interspecific pairings. Metaphycus anneckei females deposit eggs through the scale‘s ventral side and were never observed to host feed, while M. hageni females oviposit through the scale‘s dorsum and frequently host fed. The collective biological evidence agrees with morphological evidence, indicating that M. anneckei and M. hageni are discrete species. Specimens from recent collections in California and Spain were compared with specimens from earlier collections (1910s to 1980s) in Australia, California, Greece, Hawaii, Italy, Israel, Mexico, and South Africa. With the exception of Greek and Spanish material, all specimens examined were M. anneckei. We discuss the significance of this new information with respect to biological control and provide a key to separate these and other closely related black scale parasitoids.

Danzig, E.M. 2001. Mealybugs of the genera Peliococcus and Peliococcopsis from Russia and neighbouring countries (Homoptera: Coccinea: Pseudococcidae). Zoosystematica Rossica 9(1): 123-154. [Danzig2001]

Notes: The taxonomy of the genera Peliococcus Borchs. and Peliococcopsis Borchs. is discussed. Keys to species and descriptions and illustrations of most species are given. Three species are described as new: Peliococcus rosae sp. n. (Crimea and Hungary), P. loculatus sp. n. (W. Caucasus) and P. latitubulatus sp. n. (Armenia, Uzbekistan). Lectotypes are designated for 13 nominal species. The following new synonymies are established: Peliococcus balteatus (Green) = venustus (Green) = saratogensis (Rau); P. kimmericus (Kir.) = mesasiaticus (Borchs. & Kozarzh. = pseudozillae Borchs. = bitubulatus Borchs. = xerophilus Baz.; P. tritubulatus (Kir.) = darvasicus Num. & Baz.; P. turanicus (Kir.) = perfidiosus Borchs. = terrestris Borchs. = unispinus Borchs. & ter-Gr.; P. daganae (Bod.) = armeniacus Borchs.; Peliococcopsis priesneri (Laing) = caucasicus (Borchs.); Phenacoccus hordei (Lindeman) = Ph. slavonicus Laing = Peliococcus unitubulatus Borchs. & Ter-Gr.

Danzig, E.M. 2001a. A new mealybug of the genus Phenacoccus Signoret from Siberia (Hemiptera, Pseudococcidae). (In English; Summary In French). Revue Française d'Entomologie 23((1): 109-110. [Danzig2001a]

Notes: Description and illustration of adult female of Phenacoccus loiki, new sp. The host is Dactylis glomerata and it is compared to Phenacoccus avenae.

Danzig, E.M. 2001b (1999). Some aspects of the taxonomy of Palaearctic mealybugs (Hemiptera: Coccinea: Pseudococcidae). Entomologica 33: 166-167. [Danzig2001b]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. In the group of mealybugs possessing oral rim ducts, the genera Atrococcus Goux, Spilococcus Ferris, Chorizococcus McKenzie and Paracoccus Ezzat & McConnell are discussed. The genus Atrococcus is a small natural group of species, sharing similar morphology, the presence of black pigment and a Palaearctic distribution. The borders between the other genera are not clear, as has been mentioned by other authors. The genera Spilococcus and Chorizococcus are separated from each other by the number of cerarii (6-17 or fewer), while the genus Paracoccus is characterized by the presence of an anal lobe bar. In the fauna of the former USSR, there are 6 species: Spilococcus furcatispinus (Borchsenius), S. flavus (Borchsenius), S. expressus (Borchsenius), S. erianthi (Kiritshenko), S. herbaceus Danzig and S. halli McKenzie & Williams. These are all very similar morphologically and are associated with grasses (the other 7 known species in the genus Spilococcus live on dicotyledons). The placement of these species in genera based on the number of cerarii (which varies from 1 to 17 pairs) or on the presence of an anal lobe bar (S. expressus) is here considered to be artificial. Therefore, it is here suggested that the genus Spilococcus should be accepted in the wide sense (at least for the fauna in the Palaearctic region).

Dorn, B., Mattiacci, L., Bellotti, A.C. & Dorn, S. 2001. Host specificity and comparative foraging behaviour of Aenasius vexans and Acerophagus coccois, two endo-parasitoids of the cassava mealybug. Entomologia Experimentalis et Applicata 99(3): 331-339. [DornMaBe2001]

Notes: Two encyrtid parasitoids, Aenasius vexans Kerrich (Hymenoptera: Encyrtidae) and Acerophagus coccois Smith (Hymenoptera: Encyrtidae), were compared for their degree of dietary specialisation and the impact this has on their foraging strategies. Both parasitoid species are significant for biological control of the cassava mealybug, Phenacoccus herreni, Cox & Williams (Homoptera: Sternorrhyncha) a major Latin American pest of cassava, Manihot esculenta Crantz, an important root crop. Host acceptance and parasitism were analysed in seven mealybug species (with different levels of polyphagy) occurring in and around cassava fields. Results demonstrate that, in this ecosystem, An. vexans is a specialist for P. herreni while Ac. coccois is a generalist on the first and second trophic level. Of the seven mealybug species, P. herreni and P. madeirensis Green were the most acceptable hosts for Ac. coccois, followed by Ferrisia virgata Cockerell. Ac. coccois did not accept the other four mealybug species. The foraging and oviposition behaviour of individual parasitoids was observed in bioassays with cassava leaves infested by P. herreni. The two species used different strategies to locate their host. Aenasius vexans spent significantly more time walking and standing on an infested leaf and examined a host longer than did Ac. coccois. Acerophagus coccois, in contrast, spent more time for oviposition. As a consequence An. vexans parasitised more hosts in a given time than did Ac. coccois. Because the rate of offspring production of the two species did not differ, we conclude that the gregarious Ac. coccois's strategy to deposit several eggs at once might compensate for its relatively low number of ovipositions, compared with the solitary An. vexans. These findings suggest that, given the advantages and limitations of each species, a multi-species approach to biological control of P. herreni may yield best results.

Dunkelblum, E., Mori, K. & Mendel, Z. 2001 (1999). Semiochemical activity of pheromones and analogues of three Matsucoccus species (Hemiptera: Coccoidea: Matsucoccidae). Entomologica 33(1999): 391-396. [DunkelMoMe2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The sex pheromone of the Israeli pine bast scale, Matsucoccus josephi Bodenheimer & Harpaz (Homoptera: Matsucoccidae), was identified as the ketone (2E,5R,6E,8E)-5,7-dimethyl-2,6,8-decatrien-4-one. The chiral diene chain is common also to the sex pheromones of M. matsumurae Bean & Godwin and M. feytaudi Ducasse. The species-specificity of the three pheromones is due to the differences in the second side chain of these ketones. Field and GC-EAD studies indicated that the sex pheromones of each of the three Matsucoccus spp. is a potent kairomone of both males and females of the predator Elatophilus hebraicus Pericart (Hemiptera: Anthocoridae). The response of E. hebraicus to the sex pheromones of M. matsumurae and M. feytaudi is particularly interesting since it does not occur in the distribution area of these two congeneric spp. These results prompted us to prepare a series of analogues with variations in the two side chains in order to probe the structure-activity relationship of the pheromonal/kairomonal attractancy of M. josephi and E. hebraicus. Field results indicate that alterations in the common diene moiety affected the kairomonal activity, while structural changes in the second side chain markedly reduced the pheromonal activity.

Eastwell, K.C. & Bernardy, M.G. 2001. Partial characterization of a closterovirus associated with apple mealybug-transmitted little cherry disease in North America. Phytopathology 91(3): 268-273. [EastweBe2001]

Notes: Little cherry disease (LChD) is a serious economic problem of sweet cherry production in western North America where apple mealybug is the principle vector. LChD is associated with a distinct species of double-stranded (ds) RNA. In this study, filamentous virus particles were purified from LChD-infected trees and shown to contain single-stranded RNA corresponding to the previously reported dsRNA isolated from infected trees. The virus particles were characterized and were similar to monopartite members of the genus Closterovirus. A portion of the genome was sequenced and found to be most closely related to the RNA-dependent RNA polymerase of Grapevine leafroll-associated virus-3, a mealybug-transmitted closterovirus. The characteristics of the mealybug-transmitted little cherry virus in North America are very different from those of a closterovirus associated with a similar disease in Europe.

Elkins, R.B., Van Steenwyk, R.A., Varela, L.G. & Pickel, C. 2000 (1999). Pests of pear. UC Pest Management Guidelines [ElkinsVaVa2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.pears.html]. This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of pear in the U.S., including Epidiaspis leperii, Pseudococcus maritimus, P. obscurus and Quadraspidiotus perniciosus.

Elwan, A.A. 2000. Survey of the insect and mite pests associated with date palm trees in Al-Dakhliya region, Sultanate of Oman. Egyptian Journal of Agricultural Research 78(2): 653-664. [Elwan2000]

Notes: The insect pests and mites associated with infested roots, stems, leaves, infloresecence and dates [Phoenix dactylifera] of date palm trees in the Al-Dakhliya region in Oman were surveyed. The insect pests and mites associated with infested roots, stems, leaves, infloresecence and dates [Phoenix dactylifera] of date palm trees in the Al-Dakhliya region in Oman were surveyed from January 1994 to December 1996. Insect pests recorded included Aonidiella orientalis, Fiorinia linderae, Maconellicoccus hirsutus, Parlatoria blanchardii and Phoenicococcus marlatii.

Erkiliç, L.B. & Uygun, N. 2001 (1999). Observations on the population development of Pseudaulacaspis pentagona (Hemiptera: Coccoidea: Diaspididae) under semi-field conditions in the east Mediterranean region of Turkey. Entomologica 33(1999): 389. [ErkiliUy2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. Populations of Pseudaulacaspis pentagona (Targioni Tozzetti) were observed on naturally infested young peach trees (cv. 'June Gold'), kept in 2x2x2m cages covered in cheese cloth under semi-field conditions. Overwintering adult females were marked (1x2.5cm) on the plants with special glue (Tangle Trap). Both the number of crawlers produced by each female and the duration of each development stage were recorded. In each generation, 35 females were observed. P. pentagona had four generations a year and the first crawlers appeared from overwintered females at the beginning of April. The first generation was completed in May, the second in July, the third in August and the fourth in September. The duration of each generation was found to vary significantly in relation to temperature, with the first generation taking the longest (71.6 days) and the second generation the shortest (61.2 days). The number of crawlers produced by each generation was also found to vary significantly. The greatest number of crawlers (99.4 crawlers per female) was produced by the overwintering females and the lowest number of crawlers (17.9 crawlers per female) by the fourth generation.

Erler, F. & Tunç, I. 2001. A survey (1992-1996) of natural enemies of Diaspididae species in Antalya, Turkey. Phytoparasitica 29(4): 299-305. [ErlerTu2001]

Notes: 26 species of predators and 11 species of parasitoids associated with 11 species of scale insects. Hosts include Aonidiella aurantii, Lepidosaphes ulmi, Parlatoria oleae and Pseudaulacaspis pentagona.

Fang, Z.G., Wu, S.A. & Xu, H.C. 2001. A list of bamboo scale insects in China. (Homoptera: Coccoidea). (In Chinese; Summary In English). Journal of Zhejiang Forestry College 18(1): 102-110. [FangWuXu2001]

Notes: A total of 127 species of scale insects (Homoptera: Coccoidea) attacking bamboos in China are listed, including Pseudococcidae (17 genera), Coccidae (3), Aclerdidae (2), Eriococcidae (1), Asterolecaniidae (5) and Diaspididae (12 genera). Data on their distribution, hosts and the part of the bamboo that they attack are given. Two new combinations and two new synonyms are established: Planococcus bambusicola, comb. nov., was transferred from Planococcoides; Saccharicoccus bambusus, comb. nov., transferred from Palmicultor; Nipponaclerda triumpha and Bambusaspis nematosphaerica are considered to be synonyms of N. leptodemis and B. qingyuanensis, respectively.

Ferraro, M., Buglia, G.L. & Romano, F. 2001. Involvement of histone H4 acetylation in the epigenetic inheritance of different activity states of maternally and paternally derived genomes in the mealybug Planococcus citri. Chromosoma. Berlin 110(2): 93-101. [FerrarBuRo2001]

Notes: Modification of histones by acetylation is a well-known mechanism for the establishment and maintenance of specific chromatin structures with different activity states. In Planococcus citri males the paternal genome, early in development, becomes mostly inactive and heterochromatic. As we had not found methylation in the genome of P. citri, we analyzed the acetylation state of histone H4. We report here that, in males, differences in the level of histone H4 acetylation are indeed present in the two genomes of different parental origin; these differences were confirmed by treatment with the histone deacetylase inhibitor Trichostatin A. There is also evidence of acetylation of histone H4 on metaphase chromosomes. Our data therefore suggest a role of histone H4 acetylation in the imprinting of the paternal genome in P. citri males, thus supporting a role of modification of chromatin-related structural proteins in the epigenetic transmission of imprinting.

Ferrer, F. 2001. Biological control of agricultural insect pests in Venezuela; advances, achievements, and future perspectives. Biocontrol News and Information 22(3): 67N-74N. [Ferrer2001]

Notes: Biological control has been practised in Venezuela from the beginning of the 20th century, beginning with the classical introductions of Rodolia cardinalis for controlling Icerya purchasi, Aphelinus mali for the woolly apple aphid (Eriosoma lanigerum), and Apanteles thurberiae for the cotton pest Sacadodes pyralis.

Foldi, I. 2001. [List of scale insects of France (Hemiptera, Coccoidea).] Liste des cochenilles de France (Hemiptera, Coccoidea). (In French; Summary In English). Bulletin de la Société Entomologique de France 106(3): 303-308. [Foldi2001]

Notes: 381 species listed belong to 143 genera and 13 families (including Pseudococcidae, Diaspididae, Coccidae, Eriococcidae, Margarodidae, Kermesidae, Ortheziidae, Asterolecaniidae, Cerococcidae, Aclerdidae, Dactylopiidae, Lecanodiaspididae and Phoenicococcidae). A separate list of scale insects that damage cultivated and ornamental plants is also provided.

Foldi, I. & Ben-Dov, Y. 2001. A nomenclatural note on Steatococcus anonae Newstead" (Hem., Coccoidea, Margarodidae). Bulletin de la Société Entomologique de France 106(5): 449-461. [FoldiBe2001]

Notes: This note mentions the absence of the publication of the name Steatococcus anonae by Newstead. The authors assume that Newstead provided the binomen S. anonae to Varyssiere, who used it.

Foldi, I. & Watson, G.W. 2001. A new pest scale insect on avocado trees in Colombia, Laurencella colombiana, sp n. (Hemiptera: Coccoidea: Margarodidae). Annales de la Société Entomologique de France 37(3): 367-374. [FoldiWa2001]

Notes: A new scale insect pest of avocado trees (Persea americana, Lauraceae) in Colombia is becoming increasingly important at altitudes over 2000 meters above sea level, killing branches by excessive extraction of plant sap and reducing productivity by causing development of sooty mould on leaves and fruit. The adult male and female of Laurencella colombiana, sp. n. are described and illustrated. The female is recognizable by the presence of numerous distinctive cribriform cicatrices on the venter of the abdomen only. The genus Laurencella is redefined and placed in tribe Llaveiini (Monophlebinae). An identification key to the genera of Llaveiini and species of the genus Laurencella is provided. Other species of scale insect (Coccoidea) recorded on avocado worldwide are listed.

Foldi, I., Kozár, F. & Hodgson, C. 2001. Rhodococcus luberonensis, a new species of soft scale from France (Hemiptera, Coccidae). (In English; Summary In French). Bulletin de la Société Entomologique de France 106(6): 449-461. [FoldiKoHo2001]

Notes: The adults male and female, 2nd-instars male and female and pupa of a new species of Rhodococcus, R. luberonensis n. sp. (Coccidae; Coccoidea) off Rhamnus saxatilis from the south of France (Petit Lubéron mountain, Vaucluse) are described and illustrated. This new species of Rhodoccus brings the number of species in this genus to 7. The similarity of R. luberonensis to species of Eulecanium is discussed.

Follett, P.A. 2000. Arthropod pests of papaya in Hawaii. Chronica Horticulturae 40(3): 7-10. [Follet2000]

Notes: Notes are provided on the biology, harmfulness and control of the main pests of pawpaw in Hawaii. These are Tephritidae, Pseudaulacaspis pentagona, Empoasca spp. and Aphididae. Some minor pests are also listed.

Fontana, P. & Pellizzari, G. 2001 (1999). On the identity of the genus Lecanopsis Targioni Tozzetti. Entomologica 33(1999): 67-72. [FontanPe2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The genus Lecanopsis Targioni Tozzetti was generally accepted by coccidologists until 1994. In that year, on the basis that the exact facies of the genus was not known and that there was no type material, it was proposed that all the species previously included in Lecanopsis be transferred to the genus Paralecanopsis (synonymised with Lecanopsis in 1980) and that only the type species, L. rhyzophila Targioni Tozzetti, be retained in Lecanopsis, thus allowing a proper diagnosis of this group. In order to clarify the identity of the genus Lecanopsis, we have carefully checked the original descriptions of the genus and of its type species by Targioni Tozzetti and by Signoret. This work has highlighted some small mistakes in the translation of the original description from Italian or Latin to French and also some omissions, and these could have led to the conclusion that the real facies of this genus was not known and that the type species, L. rhyzophila, could not be congeneric with the other species currently included in Lecanopsis. However, some original drawings of Lecanopsis by Targioni Tozzetti, which he sent to Signoret in 1872, have been discovered in the Museum Nationale d'Histoire Naturelle, Paris. On the basis of this new information, and with the support of the authoritative opinion of two members of the International Commission of Zoological Nomenclature, we consider that (i) the genus Lecanopsis is a valid genus and propose (ii) that the species of Lecanopsis recently transferred to the genus Paralecanopsis Bodenheimer be re-assigned to the genus Lecanopsis Targioni Tozzetti.

Food and Agriculture Organization/World Health Organization Expert Committee on Food Additives 2001. Evaluation of certain food additives and contaminants. World Health Organization Technical Report Series 901: i-viii, x, 1-107. [FAO2001]

Notes: This report represents the conclusions of a Joint FAO/WHO Expert Committee convened to evaluate the safety of various food additives and contaminants, with a view to recommending Acceptable Daily Intakes (ADIs) and tolerable intakes, respectively, and to prepare specifications for the identity and purity of food additives. The first part of the report contains a general discussion of the principles governing the toxicological evaluation of food additives and contaminants (including flavouring agents), and the establishment and revision of specifications. A summary follows of the Committee's evaluations of toxicological data on various specific food additives (including cochineal extract), flavouring agents and contaminants (cadmium and tin), and of intake data on calcium from calcium salts of food additives.

Foxcroft, L.C. & Hoffmann, J.H. 2000. Dispersal of Dactylopius opuntiae Cockerell (Homoptera: Dactylopiidae), a biological control agent of Opuntia stricta (Haworth.) Haworth. (Cactaceae) in the Kruger National Park. Koedoe 43(2): 1-5. [FoxcroHo2000]

Notes: Chemical control efforts, the introduction of Cactoblastis cactorum and attempted releases of Dactylopius opuntiae Cockerell into the expanding infestation of Opuntia stricta in the Skukuza region of the Kruger National Park (KNP) have had limited success in preventing the spread and densification of O. stricta. To boost the biological control component, a new strain of D. opuntiae was introduced into KNP during 1997. The new strain established readily and has destroyed large clumps of plants in the vicinity of the release site. A large-scale redistribution programme with D. opuntiae is now needed to exploit this biological control agent to the full. In order to match the frequency of manual releases with the natural rates of spread of the insects, surveys were conducted under field conditions to determine the dispersal abilities of D. opuntiae, with regard to rate and direction of movement. Dispersal of D. opuntiae was found to be slow and restricted and that the insects need to be redistributed by placing them onto plants at approximately 10 m intervals to ensure that they become quickly and evenly distributed on the weed. This information will be crucial in the revision of the integrated management plan for O. stricta in the KNP, in integrating the cochineal and other control mechanisms.

Fraguas, J.C., Soria, S.J. & Hochmüller, D.P. 2000. [Field observations on the inter-relationships between population levels of the Brazilian ground pearl Eurhizococcus brasiliensis (Homoptera, Margarodidae) and some soil fertility factors.] (In Portuguese; Summary In English). Entomología y Vectores 7(4): 339-354. [FraguaSoHo2000]

Notes: [Original title: Observações de campo sobre as interrelações entre níveis populacionais da pérola-da-terra, Eurhizococcus brasiliensis (Homoptera, Margarodidae) e alguns fatores de fertilidade do solo.] This species is a very aggressive scale insect pest parasitizing the root system of vineyards and inducing premature dieback of fruit plants in the south of Brazil. This paper reports on information regarding the degree of influence of some soil fertility factors in the face of the magnitude of the population levels of this species in naturally infested vineyards.

Franco, J.C. & Marotta, S. 2001 (1999). A survey of mealybugs (Hemiptera: Coccoidea: Pseudococcidae) in citrus groves in continental Portugal. Entomologica 33(1999): 191-196. [FrancoMa2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] A survey of mealybugs living in citrus groves was carried out in order to elucidate the identity and relative importance of the species associated with this crop in Continental Portugal. Samples were collected in 76 citrus groves from the districts of Beja, Coimbra, Évora, Faro, Lisboa, Porto, Santarém, Setúbal and Viseu. The presence of mealybugs was detected in 93% of the citrus groves studied. Four species were identified, i.e. Planococcus citri (Risso), Pseudococcus calceolariae (Maskell), P. viburni (Signoret) and P. longispinus (Targioni Tozzetti). P. citri and P. calceolariae were the most frequent species, present in 66% and 63% of the citrus groves, respectively. P. viburni and P. longispinus were identified in 29% and 14% of the sites, respectively. Apparently, P. calceolariae has the widest geographical distribution on citrus in Portugal. The co-existence of at least two species in the same citrus grove was found in 59% of the cases. P. citri and P. calceolariae were found associated in 37% of the citrus groves; P. viburni was always found associated with at least one of the three other species.

Franco, J.C., Silva, B.E. & Carvalho, J.P. 2000. (In Portuguese). In: , [Mealybugs associated with citrus in Portugal.] Cochonilhas-algodão (Hemiptera, Pseudococcidae) associadas aos citrinos em Portugal. ISA Press, Lisbon. 142 pp. [FrancoSiCa2000]

Notes: Synonymy, common names, morphology, geographic distribution, hosts, life cycles and biology are topics covered.

Franke, I.H., Fegan, M., Hayward, C., Leonard, G. & Sly, L.I. 2000. Molecular detection of Gluconacetobacter sacchari associated with the pink sugarcane mealybug Saccharicoccus sacchari (Cockerell) and the sugarcane leaf sheath microenvironment by FISH and PCR. FEMS Microbiology Ecology 31(1): 61-71. [FrankeFeHa2000]

Notes: Molecular tools for the detection of the newly described acetic acid bacterium Gluconacetobacter sacchari from the pink sugarcane mealybug, Saccharicoccus sacchari Cockerell (Homoptera: Pseudococcidae), and in the sugarcane leaf sheath microenvironment were developed. G. sacchari specific 16S rRNA-targeted oligonucleotide primers were designed and used in PCR amplification of G. sacchari DNA directly from mealybugs, and in a nested PCR to detect low numbers of the bacteria from sugarcane leaf sheath fluid and cane internode scrapings. A sensitivity level of detection of 40-400 cells/reaction was obtained using PCR from exponentially grown bacterial cultures and of 1-10 cells in cane internode scrapings and leaf sheath fluid samples using nested PCR. The specificity of the primer set was demonstrated by the lack of amplification product formation in PCR by closely related acetic acid bacteria, including Gluconacetobacter liquefaciens, and Gluconacetobacter diazotrophicus. A Cy3 labeled probe for G. sacchari was designed and shown to be specific for the species. Investigation of the mealybug microenvironment by whole cell fluorescent in situ hybridization revealed that G. sacchari appears to represent only a minor proportion of the population of the microbiota in the mealybugs tested. This study has shown the usefulness of 16S rRNA-based molecular tools in the identification and detection of G. sacchari from environmental samples and will allow these tools to be used in further ecological research.

Fukatsu, T. & Nikoh, N. 2000. Endosymbiotic microbiota of the bamboo pseudococcid Antonina crawii (Insecta, Homoptera). Applied and Environmental Microbiology 66(2): 643-650. [FukatsNi2000]

Notes: The intracellular symbiotic microbiota of the bamboo pseudococcid Antonina crawii was characterized by performing a molecular phylogenetic analysis in combination with in situ hybridization. Almost the entire length of the bacterial 16S rRNA gene was amplified and cloned from A. crawii whole DNA. Restriction fragment length polymorphism analysis revealed that the clones obtained included three distinct types of sequences. Nucleotide sequences of the three types were determined and subjected to a molecular phylogenetic analysis. The first sequence was a member of the gamma subdivision of the division Proteobacteria (gamma-Proteobacteria) to which no sequences in the database were closely related, although the sequences of endosymbionts of other homopterans, such as psyllids and aphids, were distantly related. The second sequence was a beta-Proteobacteria sequence and formed a monophyletic group with the sequences of endosymbionts from other pseudococcids. The third sequence exhibited a high level of similarity to sequences of Spiroplasma spp. from ladybird beetles and a tick. Localization of the endosymbionts was determined by using tissue sections of A. crawii and in situ hybridization with specific oligonucleotide probes. The gamma- and beta-Proteobacteria symbionts were packed in the cytoplasm of the same mycetocytes (or bacteriocytes) and formed a large mycetome (or bacteriome) in the abdomen. The spiroplasma symbionts were also present intracellularly in various tissues at a low density. We observed that the anterior poles of developing eggs in the ovaries were infected by the gamma- and beta-Proteobacteria symbionts in a systematic way which ensured vertical transmission. Five representative pseudococcids were examined by performing diagnostic PCR experiments with specific primers; the beta-Proteobacteria symbiont was detected in all five pseudococcids, the gamma-Proteobacteria symbiont was found in three, and the spiroplasma symbiont was detected only in A. crawii.

Gaimari, S.D. & Tanasijtshuk, V.N. 2001. A new leucopine genus (Diptera: Chamaemyiidae) with species attacking Ceroplastes wax scales (Hemiptera: Coccidae) in South America. Systematic Entomology 26(3): 311-328. [GaimarTa2001]

Notes: A new genus, Echinoleucopis, is proposed within Chamaemyiidae (Diptera). All known species of the genus are from South America and are predators on eggs within an ovisac of wax scales in genus Ceroplastes Gray (Hemiptera: Coccidae).

Ganeshan, S. 2000. Biological control of sugarcane pests in Mauritius: current status and future prospects. 3-9. In: Allsopp, P.G. (Ed.) & Suasa Ard, W., Sugarcane Pest Management in the New Millennium. 4th Sugarcane Entomology Workshop International Society of Sugar Cane. International Society of Sugar Cane Technologists, Indooroopilly, Australia. [Ganesh2000]

Notes: Manipulation of the environment through classical biological control and adoption of cultural measures, such as weed control and cane trashing, have been the main pest management strategies implemented for many years in Mauritius. Several species of parasitoids and predators have been introduced and released against major pests including Aulacaspis tegalensis (Zehntner). A. tegalensis remains a chronic pest causing severe but localised losses in some regions. Outbreaks of occasional pests such as the soft scale Saccharipulvinaria iceryi (Signoret) have been managed by implementation of measures favouring the multiplication of its natural enemies. However, studies on the economic importance of these pests and the presence of several species of parasitoids have revealed that their control by insecticides might not be cost effective and could be detrimental to the parasitoids.

Gang, F.Z. & Jun, L. 2000. [Biological observations and control of Kuwanaspis howardi] (In Chinese; Summary In English). Journal of Bamboo Research 19(2): 78-80. [GangJu2000]

Notes: Kuwanaspis howardi is mainly a sucking pest of bamboo shoots of Phyllostachys praecox, P. dulcis, and other [?Phyllostachys] species in China. The paper gives a brief account of its biology, life cycle and control. Observations in [Zhejiang Province] have shown that there are two generations per year, with overwintering by the fertilized adult female. Effective control of the pest is achieved either by injection of omethoate (40%) at the rate of 1:10 in the bamboos chamber [culm] or by smearing on the surface of the bamboo. Some control can be achieved using ordinary washing-powder (1:200) puffed onto the bamboo straw.

Gardosik, S.M. 2001. Aspidiotus cryptomeriae Kuwana, an armored scale pest of conifers (Homoptera: Diaspididae). Regulatory Horticulture (PA Dept. of Agric.) 27(200): 23-25. [Gardos2001]

Notes: Hosts and distribution, identification, life history, damage and control are discussed.

Geiger, C.A., Daane, K.M., Bentley, W.J., Yokota, G.Y. & Martin, L.A. 2001. Sampling program for grape mealybugs improves pest management. California Agriculture 55(3): 19-27. [GeigerDaBe2001]

Notes: The results of a mealybug study in Central Valley vineyards, designed to develop sampling guidelines, reveal that mealybug distribution on vines varies greatly through the season and that mealybugs usually prefer concealed locations, such as under bark. This combination makes sampling difficult. A number of sampling techniques were compared. Three-or 5-minute timed counts were most efficient because samplers could follow the mealybugs' movement over the season. Midseason counts were much better predictors of damage at harvest than early season counts. This research confirms past control guidelines and opens new control options. Grape bunches touching vine trunks or spurs will have higher damage. Removing these bunches or using barriers between bunches and mealybug oviposition sites can also reduce damage.

Geraud-Pouey, F., Chirinos, D.T. & Romay, G. 2001. [Physical effect of exfoliation of guava tree bark on Capulinia sp. near to jaboticabae von Ihering (Hem.: Eriococcidae)] Efecto fisico de las exfoliaciones de la corteza del guayabo (Psidium guajava) sobre Capulinia sp. cercana a jaboticabae. (In Spanish; Summary In English). Entomotropica 16(1): 21-27. [GeraudChRo2001]

Notes: Since it appeared in Venezuela in early 1993, the guava cottony scale, Capulinia sp. has become the greatest entomological problem on guava trees, Psidium guajava. However, in the field, trees with smooth bark (with little or without exfoliations) tend to be much less infested by this insect. During May-July 1999, the physical effects of bark exfoliation on the establishment, survival and development of Capulinia sp. was assessed, in the laboratory. Three host plant conditions were included: with natural exfoliations; with bark mechanically smoothed (smooth bark); and exfoliations substituted by pieces of absorbent paper adhered to the smoothed bark (artificial exfoliations). Bark smoothness could constitute a source of resistance of guava plants to diminish the incidence and damages by Capulinia sp.

Gertsson, C.A. 2001. An annotated checklist of the scale insects (Homoptera: Coccoidea) of Sweden. [Förtechnung över Sveriges sköldlöss.] (In English; Summary In Swedish). Entomologisk Tidskrift. Stockholm 122(3): 123-130. [Gertss2001]

Notes: This checklist identifies 72 species records from the 30 Swedish provinces. 13 additional species are listed from Norway, Finland and/or Denmark. Remarks are given for 57 species, including notes on records and references to the literature. A bibliography is concluded.

Ghabbour, M.W. 2001 (1999). Descriptions of the first-instar nymphs of three species of Lepidosaphes Shimer and a species of Insulaspis Mamet (Hemiptera: Diaspididae). Entomologica 33(1999): 73-83. [Ghabbo2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The first-instar nymphs of two species of the genus Lepidosaphes (namely L. beckii (Newman) and L. ficus (Signoret)) are described and illustrated and compared with the first-instar nymph of L. ulmi. The first-instar nymph of Insulaspis tapleyi (Williams) is also described and illustrated, and the differences between I. tapleyi and I. pallidula (Green) are discussed. Keys are provided for the separation of the first-instar nymphs of these five species.

Ghosh, A.B. & Ghose, S.K. 2000. Effect of Some Ecological Factors on the Nymphal Development of Mealybug, Nipaecoccus viridis (Newstead) Pseudococcidae : Hemiptera). Environment & Ecology 18(4): 922-925. [GhoshGh2000]

Notes: The developmental periods of all female nymphs and up to second male nymphs of the mealybug, Nipaecoccus viridis (Newstead), were prolonged when reared individually. The development of female and male nymphs were also delayed at constant relative humidity of 40, 60 and 80%. Third nymphal females at 80% and fourth nymphal males at 60 and 80% moulted 5 days later than at other humidity levels. The constant exposure to light had induced moulting earlier than darkness in second nymphal females only. The presence of attending ant, Camponotus oblongus Sm. accelerated the development and moulting in first instar nymph.

Giliomee, J.H. 2001 (1999). The Asterolecaniidae (Hemiptera: Coccoidea) of South Africa. Entomologica 33(1999): 85-89. [Giliom2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The 16 species of Asterolecaniidae known from South Africa can be tentatively placed in six genera based on the characters of the adult female (Abditococcus, Asterodiaspis, Asterolecanium, Bambusaspis, Planchonia and Russellaspis. Some species do not fit into these genera as presently defined, requiring a redefinition of some genera or the creation of new ones. However, it appears that the characters on which the genera of the Asterolecaniidae are presently based are too few in number and unreliable, making a detailed study of other characters of the female and eight other stages necessary for a better understanding of the relationships within this family.

Gillani, W.A. & Copland, M.J.W. 2001 (1999). Defensive behaviour of the longtailed mealybug Pseudococcus longispinus (Targioni Tozzetti) (Hemiptera: Pseudococcidae) against the brown lacewing Sympherobius fallax Navas (Neuroptera: Hemerobiidae). Entomologica 33(1999): 279-285. [GillanCo2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The defensive tactics of 2nd- and 3rd-instar nymphal and adult Pseudococcus longispinus (Targioni Tozzetti) against larval Sympherobius fallax Navas were studied. When attacked by 1st- and 2nd-instar predators, these three mealybug stages were able to secrete ostiolar fluids which spread and hardened on the mouthparts of the predators; 1st-instar mealybugs appeared to be unable to secrete ostiolar fluids. Adult mealybugs also successfully defended themselves against 1st-instar predators by simply pushing them away with their cerarial wax fringe. Of the lacewing stages, 3rd-instar S. fallax larvae were the most successful at overcoming the defense tactics of P. longispinus.

Gilrein, D. 2001. Tipping the scales. Grounds Maintenance 36(9): 10,14-16. [Gilrei2001]

Notes: Types of scales discussed in a general way include mealybugs (Pseudococcidae), soft scales, wax scales and tortoise scales (Coccidae), armored scales (Diaspididae), felt scales or eriococcids (Eriococcidae), margarodid scales (Margarodidae), bark crevice scales (Cryptococcidae), gall scales or kermesids (Kermesidae) and pit scales or asterolecaniids (Asterolecaniidae). Description of damage, hosts and control are discussed.

Gitirana Neto, J., Carvalho, C.F., Souza, B. & Santa Cecilia, L.V.C. 2000. [Population dynamics of the Pinnaspis aspidistrae (Signoret, 1869) (Hemiptera: Diaspididae) in citrus trees in Lavras - MG.] (In Portuguese; Summary In English). Ciencia e Agrotecnologia 24(3): 632-645. [GitiraCaSo2000]

Notes: [Original title: Flutuaçao populacional de Pinnaspis aspidistrae (Signoret, 1869) (Hemiptera: Diaspididae) em citros, na regiao de Lavras - MG.] The population dynamics of Pinnaspis aspidistrae was studied from May 1992 to April 1996 in a citrus orchard located in Lavras, Minas Gerais, Brazil. Leaves, twigs and fruits infested with male second instar nymphs were collected weekly from 5 trees of each citrus cultivar (Natal, Valencia, Baia and Ponkan). P. aspidistrae was found all over the orchard in all months sampled, with infestations varying in different hosts and seasons. Generally, the greatest infestations occurred in periods characterized by low temperature and rainfall. The highest populations were observed during September/October for Natal, during August for Valencia and during May/August for Baia, while Ponkan exhibited low population densities throughout the study.

Gomez, L.D. & Kisimova-Horovitz, L. 2001. A new species of Septobasidium from Costa Rica. Mycotaxon 80: 255-259. [GomezKi2001]

Notes: Septobasidium wilsonianum associated with scale insects (Diaspididae, Coccoidea, Homoptera) attached to Topobea multiflora, Melastomataceae, is described and illustrated as a new species. It is the first report of a species of Septobasidium occurring on a member of the Melastomataceae.

Goncalves-Gevasio, R.D.R. & Santa-Cecilia, L.V.C. 2001. Food consumption of Chrysoperla externa in different developmental phases of Dysmicoccus brevipes, under laboratory conditions. (In Portuguese). Pesquisa Agropecuaria Brasileira 36(2): 387-391. [GoncalSa2001]

Notes: The food consumption and preference of larvae of Chrysoperla externa Hagen, 1861 Neuroptera: Chrysopidae) larval were studied in laboratory under 25 +/-2 degrees C, 70 +/- 10% RH and 12 hours photophase, utilizing the pineapple mealybug (Dysmicoccus brevipes Cockerell, 1893, Hemiptera: Pseudococcidae) as a prey. The results showed that the predator's larval phases consumed on average of 70, 50 and 15,8 mealybugs of the first, second and third instar, respectively, and 10 adult females amounting to 145,8 individuals. It was noticed that 5 1% of the total number of mealybugs were consumed during the predator third instar. The high number of mealybugs consumed shows that C. externa presents a potential to be utilized in biological control of the mealybug D. brevipes.

Gonsalves, D. 2000. Grapevine leafroll-associated closteroviruses: Characterization and transgenic resistance. Phytopathology 90(6)(Suppl.): S91. [Gonsal2000]

Notes: GLRaV 3 is transmitted by mealybugs, while GLRaV 1 may be transmitted by scale insects.

Gonzalez, P. 2000. First record of the genera Chilecoccus and Icelococcus, and two Acanthococcus species (Hemiptera: Eriococcidae) for Argentina. Revista de la Sociedad Entomologica Argentina 59(1-4): 51-52. [Gonzal2000]

Notes: Two genera: Icelococcus Miller & Gonzalez and Chilecoccus Miller & Gonzalez, and two Acanthococcus species are listed on Nothofagus (Fagaceae) for the first time in Argentina in this paper. They were also cited for Chile on the same hosts.

González, R.H., Poblete G., J. & Barría P., G. 2001. [Bionomics of the tree fruit mealybug in Chile, Pseudococcus viburni(Signoret).] El chanchito blanco de los fruitales en Chile, Pseudococcus viburni (Signoret), (Homoptera: Pseudococcidae). (In Spanish; Summary In English). Revista Fruticola 22(1): 17-26. [GonzalPoBa2001]

Gonçalves Gervasio, R, de C.R. & Santa Cecilia, L.V.C. 2001. [Food consumption of Chrysoperla externa in different developmental phases of Dysmicoccus brevipes, under laboratory conditions.] (In Portuguese; Summary In English). Pesquisa Agropecuaria Brasileira 3(2): 387-391. [GoncalDe2001]

Notes: The food consumption and preference of Chrysoperla externa larvae were studied in a laboratory under 25+̊C, 70+10% RH and 12 h photophase, utilizing the pineapple mealybug (Dysmicoccus brevipes) as a prey. The high number of mealybugs consumed indicated that C. externa is a potential biological control agent of D.brevipes.

Grafton-Cardwell, E.E., Morse, J.G., O'Connell, N.V. & Phillips, P.A. 2000. Pests of citrus. UC Pest Management Guidelines [GraftoMoOC2000]

Notes: [http://www.ipm.ucdavis.edu/PMG/selectnewpest.citrus.html This update to the University of California IPM Pest Management Guidelines contains descriptions and control recommendations for insects, arthropod and nematode diseases of citrus in the U.S., including Aonidiella aurantii, A. citrina, Coccus hesperidum, C. pseudomagoliarum, Icerya purchasi, Lepidosaphes beckii and Saissetia oleae.

Grafton-Cardwell, E.E., Ouyang, Y., Striggow, R. & Vehrs, S. 2001. Armored scale insecticide resistance challenge San Joaquin Valley citrus growers. California Agriculture 55(5): 20-25. [GraftoOuSt2001]

Notes: Aonidiella aurantii and A. citrina are among the insect pests discussed.

Gross, S., Biraty, Y. & Gal, S. 2001. [Using powdery and microcapsular preparations to decimate ant populations on citrus trees.] (In Hebrew; Summary In English). Alon Hanotea 55(5): 219-221. [GrossBiGa2001]

Notes: Citrus trees were treated with chlorpyrifos (5% powder formulation at 100 g/tree, or 1 and 2% microcapsule formulation sprayed on and around the trunk, at 1 litre/tree) or carbosulfan (2% powder formulation applied on the trunk, 100 g/tree) to study their effects against ant infestation. All treatments were effective in controlling the ant population. Also, low mealybug fruit infestation and higher exportable yield were recorded from the treated plants compared with the untreated controls.

Gross, S., Gefen, D., Rotman, N., Tadmor, U., Zemer, B., Gotlib, A. & Gefen, Y. 2000. Chemical control of the spherical mealybug (Nipaecoccus viridis) (Newstead) in citrus. (In Hebrew; Summary In English). Alon Hanotea 54(6): 234-240. [GrossGeRo2000]

Notes: The spherical mealybug infests all parts of the citrus tree. Its damage is expressed by excretion of large quantities of honeydew which encourages sooty mould to cover most parts of the tree, appearance of irregular green spots on the fruit and malformed fruit which are rendered inexportable. Herewith we report on field trials, which compared the efficacy of various chemical treatments to control the spherical mealybug. Identical dosages of Mospilan (acetamiprid 20 SP) were applied at two stages of development: before blossom and on small fruits (15-20 mm). Those treatments were compared to the standard chemical treatment of chlorpyrifos, and to non-treated control plots. The results indicate that there is no advantage in applying Mospilan prior to blossom. Effective control was achieved by Mospilan 0.05% and 0.075% sprayed on small fruits or by 0.3% chlorpyrifos at both stages of development.

Guerrieri, E. & Noyes, J.S. 2002. An unusual genus and species of Encyrtidae (Hymenoptera: Chalcidoidea) from Australia reared from soft scale insects (Hemiptera: Coccidae). Journal of Natural History 36(4): 443-448. [GuerriNo2002]

Notes: Arketypon vaderi gen. et sp. nov. is described from Australia and its systematic position is discussed. It was reared as a gregarious endoparasitoid of Myzolecanium sp. (Hemiptera: Coccidae) found in the nest of Camponotus sp. (Hymenoptera: Formicidae) nesting in a hollow stem of Sonneratia alba Smith (Sonneratiaceae).

Guerrieri, E., Pedata, P.A., Romani, R., Isidoro, N. & Bin, F. 2001. Functional anatomy of male antennal glands in three species of Encyrtidae (Hymenoptera : Chalcidoidea). Journal of Natural History 35(1): 41-54. [GuerriPeRo2001]

Notes: The occurrence of male antennal glands in three hymenopteran parasitoids belonging to the family Encyrtidae (Leptomastix dactylopii, Rhopus meridionalis and Asitus phragmitis) is reported for the first time. The ultrastructure and function of these glands are described in detail. The possible behavioural and taxonomical implications are discussed. Hosts include Chaetococcus phragmitis and Planococcus citri.

Gullan, P.J. 2001 (1999). Why the taxon Homoptera does not exist. Entomologica 33(1999): 101-104. [Gullan2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Many researchers still use the name Homoptera for the higher taxon of any member of the Sternorrhyncha (Aleyrodoidea, Aphidoidea, Coccoidea and Psylloidea) or Auchenorrhyncha (Cercopoidea, Cicadoidea, Cicadelloidea and Fulgoroidea), or to refer collectively to the Sternorrhyncha plus Auchenorrhyncha. Recent work based on morphological and molecular studies provides phylogenetic evidence that the Homoptera is paraphyletic and therefore its use should be abandoned.

Gullan, P.J. & Cook, L.G. 2001 (1999). Are cochineal insects Eriococcids? Entomologica 33(1999): 91-99. [GullanCo2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Scale insects of the genus Dactylopius Costa, which all feed on cacti, are commonly called cochineal insects. Currently there are nine described species placed in their own family, the Dactylopiidae, based on a few unique morphological features. Here we review available biological, morphological and karyotype information on Dactylopius and report on cladistic analyses of morphological data (from first-instar nymphs plus adult females, and adult males separately) and molecular data (from the nuclear gene 18S rDNA and the mitochondrial gene cytochrome oxidase II) from Dactylopius and its potential relatives. We suggest that Dactylopius belongs with the eriococcids and we discuss the nomenclatural implications of this placement.

Gullan, P.J. & Sjaarda, A.W. 2001. Trans-Tasman Platycoelostoma Morrison (Hemiptera: Coccoidea: Margarodidae) on endemic Cupressaceae, and the phylogenetic history of margarodids. Systematic Entomology 26: 257-278. [GullanSj2001]

Notes: A new species of Platycoelostoma Morrison (Hemiptera: Coccoidea: Margarodidae) is described from the Tasmanian endemic conifer, Diselma archeri (Cupressaceae), growing in alpine heathland. Its sister species, P. compressa (Maskell), occurs in alpine and subalpine New Zealand on Libocedrus bidwilli (Cupressaceae). Vicariance dating provides a minimum age for Platycoelostoma of about 80 Ma. Cladistic analysis of morphological data from adult females and first-instar nymphs suggests that Platycoeostoma is the sister genus of Callipappus Guerin-Meneville and also is more closely related to members of subfamilies Margarodinae and Xylococcinae than to the New Zealand and South American genera of Coelostomidiinae, to which Platycoelostoma traditionally has been linked. The analysis also indicates that most traditional tribal groupings within Margarodidae are monophyletic, but that three (Coelostomidiinae, Margarodinae and Xylococcinae) of the five subfamilies in the widely used higher classification of Morrison may be either paraphyletic or polyphyletic as currently defined. Only Monophlebinae and Steingeliinae have adequate support from morphological characters. Relationships among subfamilies and many tribes are unresolved. These results lend some support to the classification previously proposed by Koteja, in which all margarodid subfamilies and a few tribes are elevated to the rank of family. Furthermore, the analysis suggests that Marchalini and Monophlebinae (in which all immature female instars have well developed appendages) are derived with respect to other margarodids and thus development via legless intermediate female instars ('cyst' form) may be plesiomorphic within margarodids. This hypothesis is weakly supported and requires corroboration from independent data. Platycoelostoma is transferred to tribe Callipappini, which is redefined, but the subfamily placement of this tribe must remain uncertain until the higher classification and rank of the various margarodid taxa can be re-evaluated. A revised generic description of Platycoelostoma, based on the adult and intermediate female instars and the first instar of both species, and a key to separate the female instars of both species are provided. The first-instar nymph, all female instars and the third-instar male (the prepupa) of Platycoelostoma tasmanicum sp.n. are described and illustrated. Scanning electron micrographs depict the unusual cuticular features of the third-instar female.

Handique, P.K. & Baruah, R. 2000. [Evaluation of Biomix-1 against mealy bug (Maconellicoccus hirsutus Green) of mulberry.] Indian Journal of Sericulture 39(1): 79-80. [HandiqBa2000]

Notes: Laboratory studies showed that Biomix-1 at a 1% concentration [based on plant extracts of unstated composition] was as effective against Maconellicoccus hirsutus as dimethoate at 0.1%.

Hansen, J.D. 2001. Ultrasound treatments to control surface pests of fruit. HortTechnology 11(2): 186-188. [Hansen2001]

Notes: Durations of ultrasound treatments were evaluated for efficacy in removing or destroying external pests of apples (Malus sylvestris var. domestica). Ultrasound did not remove San Jose scale (Quadraspidiotus perniciosus, Homoptera: Diaspididae), from the fruit surface. Ultrasound, which can be incorporated in the packing line, shows promise as a postharvest phytosanitation treatment against external pests.

Hare, J.D. & Morgan, D.J.W. 2000. Chemical conspicuousness of an herbivore to its natural enemy: Effect of feeding site selection. Ecology 81(2): 509-519. [HareMo2000]

Notes: A physical refuge from the parasitoid Aphytis melinus is provided to the California red scale Aonidiella aurantii by the interior bark substrates of citrus trees, even though scales have lower fitness on bark in the absence of A. melinus. How bark-reared scales escape parasitism was unclear because A. melinus searches the interior of trees as effectively as the exterior. Host identification in A. melinus is mediated by a kairomone, O-caffeoyltyrosine, in scale covers. O-caffeoyltyrosine concentration varies with scale age and rearing conditions. We hypothesized that the reduced acceptance of bark-reared scale may be due, in part, to reduced quantities of O-caffeoyltyrosine in their covers. We reared scales on bark, leaves, and fruit of lemon and orange trees in the field at monthly intervals and then collected the scales and measured them. We bioassayed covers for their acceptability to A. melinus in the laboratory and then determined their O-caffeoyltyrosine content. Even after adjusting for the differences in scale body size, O-caffeoyltyrosine content in bark-reared scale covers was 45-85% less than that in covers of leaf- or fruit-reared scales, depending upon cultivar and rearing date. Covers of bark-reared scales were selected for probing only 40-45% of the time when compared to leaf-reared scales. Covers with the highest levels of O-caffeoyltyrosine were most likely to be selected for probing. We conclude that part of the mechanism by which California red scales avoid discovery on bark is through reduced O-caffeoyltyrosine content in their covers. This reduction is probably a consequence of the reduced nutritional quality of bark as a substrate for scale survival and growth.

Harris, P. 2001. The history of host range testing for classical weed biocontrol agents. Agriculture and Agri-Food Canada, Research Branch http://res2.agr.ca/lethbridge/weedb [Harris2001]

Notes: Dactylopius coccus is mentioned as a cactus-feeding mealybug and a commercial source of red dye.

Heckroth, H.-P., Fiala, B. & Maschwitz, U. 2001. Integration of scale insects (Hemiptera: Coccidae) in the southeast Asian ant-plant (Crematogaster (Formicidae)-Macaranga (Euphorbiaceae)) system. Entomologica 33: 287-295. [HeckroFiMa2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The coccid colonisers of myrmecophytic Macaranga were tested for their trophic integration into the mutualistic Crematogaster-Macaranga ant-plant system. Honeydew secretion by these mostly endophytic scale insects was observed in Coccus caviramicolus Morrison, C. penangensis Morrison, P. secretus Morrison, C. tumuliferus Morrison, C. tumuliferus var. C.84 and in two other Macaranga coccid species. However, the use of coccids as protein-rich food by ants under normal and starvation conditions was not observed.

Heidari, M. 2001 (1999). The intrinsic rate of increase and temperature co-efficients of the Comstock mealybug, Pseudococcus comstocki (Kuwana) (Hemiptera: Coccoidea: Pseudococcidae). Entomologica 33(1999): 297-303. [Heidar2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Life history studies on the development of Pseudococcus comstocki (Kuwana) were made under laboratory conditions at four constant temperatures, namely 18, 22, 26 and 30̊C. Temperature proved to have a significant effect on development, survival, reproduction and longevity, and on the sex ratio of P. comstocki. Reproduction, survival and longevity were greatest at 22-26̊C and lowest at 30̊C, which was detrimental to the nymphs. It was also found that the sex ratio became female biased at 30̊C. On the basis of these data, the intrinsic rate of natural increase (rm) was computed to be 0.05, 0.07, 0.10 and 0.08 at the above temperatures and the gross reproduction rate (GRR) was also temperature dependent and was calculated to be 153, 249, 210 and 57. The lower temperature threshold for development (TL) was estimated to be 11.0̊C and the time from birth to adulthood (K) 523 degree days.

Heidari, M. 2001a (1999). Influence of host-plant physical defenses on the searching behaviour and efficacy of two coccinellid predators of the obscure mealybug, Pseudococcus viburni (Signoret). Entomologica 33: 397-402. [Heidar2001a]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Laboratory and glasshouse trials were conducted to investigate the influence of host-plant physical structure on the predation of the obscure mealybug, Pseudococcus viburni (Signoret) (Coccoidea: Pseudococcidae) by two coccinellid predators: Cryptolaemus montrouzieri (Mulsant) and Nephus reunioni Fürsch. Glasshouse trials showed that good control was achieved by C. montrouzieri irrespective of the hairiness of the plant species, but that N. reunioni was more effective on smooth leaves, such as those of Citrus, coffee and Passiflora, than on hairy plants, such as Streptocarpus and tomato, on which the density of the trichomes and their sticky exudations significantly reduced the walking speed and other searching parameters. It is concluded that the size of the predator and the type of trichomes have a marked influence on the level of mealybug control and that this could influence the choice of predators.

Helal, E.M., Donia, A.R., El Hamid M.M.A. & Zakzouk, E.A. 2000. Abundance of insect and mite species in some citrus orchards. Proceedings of the International Conference on Integrated Fruit Production No. 525: 443-453. [HelalDoEl2000]

Notes: [Conference held in Leuven, Belgium, 27 July-1 August 1998.] A survey of pests and mites occurring on Citrus sinensis Qsheck var. Washington navel and Citrus aurantium L. at Alexandria and Damanhour, Egypt, was carried out for a whole year. Seven insect pests were found, of which the citrus white fly, Aleurotrachelus citri was the most common. The predator Chrysopa vulgaris [Chrysoperla carnea] also occurred, attacking the citrus mealy bug Planococcus citri. Eleven phytophagous, predaceous and saprophagous mite species prevailed on orange and sour orange in both experimental locations. In addition, four mite species were found associated with weeds growing in citrus orchards at Alexandria, including one phytophagous and three predaceous species. The results indicated that Aleurotrachelus citri is a major pest of citrus. Its pattern of distribution was followed at both locations. The correlations between temperature and relative humidity and the abundance of insects and mites on both citrus species were studied. The correlations between the predaceous mites and their prey were also investigated.

Hellemans, A. & Goossens, F. 2000. [Control of common scale (Parthenolecanium pomeranicum) on Taxus.] Bestrijding van gewone dopluis (Parthenolecanium pomeranicum) op Taxus. (In Dutch). Verbondsnieuws 44(19): 27-29. [HellemGo2000]

Notes: Studies were carried out in 1998 in Belgium to compare the efficacy of Mitac (200 g amitraz/litre), Deltanet 400 EC (400 g furathiocarb/litre), Confidor 200 SL (200 g imidacloprid/litre), Applaud (250 g buprofezin/litre), and Karate 25 WG (2.5% lambda-cyhalothrin) for the control of P. pomeranicum on 7- and 8-year-old T. baccata. Treatments were carried during the last larval stage (22 and 29 April 1998) and after ova budding (29 June and 8 July 1998). The most effective insecticides for the first period (last larval stage) were Mitac, Deltanet and Confidor, and Mitac, Deltanet and Karate for the 2nd period.

Henderson, R. 2001 (1999). The structure and function of the test of New Zealand male soft scale insects. Entomologica 33(1999): 167. [Hender2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. Male scale insects (Hemiptera: Coccoidea) undergo a metamorphosis from scale-like nymph through prepupa and pupa to winged adult. The nymphal instar preceding the prepupa secretes a protective cover or test under which these complex life changes take place. Each family of scale insects is characterised by a different type of male covering, whether cocoon, cap or test. Male mealybugs make rather fluffy cocoons from cottony wax strands, eriococcids produce woven wax covers, while armoured scales incorporate their moulted skins into waxy caps. The Coccidae or soft scales construct glassy wax tests. This test, being rigid, needs a mechanism to allow for the emergence of the adult male. A suture across the posterior third of the test enables the back plate to flex at a pair of hinges (so forming something akin to an "up-and-over garage door"). The hinges are secreted by groups of tubular ducts on the dorsum of the 2nd-instar males. The rows of hexagonal wax-plates found in the male tests of most New Zealand Coccidae are apparently unique in the way they are constructed. Scanning electron micrographs show the detail and diversity of the hinge types and how the hexagonal plate structure of the test is distinct from the back plate suture. Very little research has been published on male scale insects and even less on their coverings. In comparison with North American species, only two species of New Zealand native soft scales have male tests similar to those in other parts of the world.

Henderson, R.C. 2000. The status of Parlatoria species (Hemiptera: Coccoidea: Diaspididae) in New Zealand. New Zealand Entomologist 23: 51-53. [Hender2000]

Notes: Three species of Parlatoria are found in New Zealand: Parlatoria desolator McKenzie, P. fulleri Morrison and P. pittospori Maskell. Previous records of P. pergandii Comstock and P. virescens Maskell from New Zealand are based on misidentifications of P. desolator. The sole record of P. ziziphi (Lucas) is based on a dubious quarantine record.

Henderson, R.C. 2001a. New synonymies for two armoured scale insects (Hemiptera: Coccoidea: Diaspididae) on Dysoxylum spectabile. New Zealand Entomologist 24: 89-90. [Hender2001a]

Notes: The taxonomy of Aspidiotus nerii and Pinnaspis dysoxyli are clarified in this report.

Henderson, R.C. & Rhode, B.E. 2001. The hinged back plate mechanism in glassy wax tests of New Zealand male soft scale insects (Hemiptera : Coccoidea : Coccidae). Arthropod Structure & Development 30(1): 1-14. [HenderRh2001]

Notes: Male scale insects (Hemiptera: Coccoidea) undergo a metamorphosis of the neometabola type, from scale-like nymph through prepupa and pupa to winged adult. The nymphal instar before prepupa secretes a waxy protective covering that remains in place throughout metamorphosis and these covers are characteristic of each family of scale insects. Most scale insect families (e.g. mealybugs, eriococcids, diaspidids) have rather loosely woven male covers, but male nymphs in the family Coccidae (soft scales) construct more rigid, glassy wax tests, which need a special mechanism for adult emergence. In the New Zealand male soft scales, a suture across the posterior quarter of the test enables the back plate to flex at a pair of hinges, to be raised up off the substrate, and so allow egress. The waxy back plate hinges are secreted by groups of tubular ducts on the abdominal dorsum of 2nd-instar males, during construction of the test, scanning electron micrographs (SEMs) show the detail and diversity of hinge types. The wax tests of most New Zealand Coccidae, both female and male, are apparently unique in that they are constructed in rows of hexagonal plates, separated by sutures, however in the male test, the sutures are all fused except for the back plate suture. The two species in the endemic New Zealand genus Pounamococcus have male tests more like those of species in the Australian genus Austrolecanium.

Hickel, E.R., Peruzzo, E.L. & Schuck, E. 2001. Control of the Ground-Pearl Eurhizococcus brasiliensis (Hempel) (Homoptera: Margarodidae) with chemigation. Neotropical Entomology 30(1): 125-132. [HickelPeSc2001]

Notes: The ground-pearl Eurhizococcus brasiliensis (Hempel) (Homoptera: Margarodidae) is the main pest of vineyards in Southern Brazil. The normal pest control strategies do not affect the pest population because the insects are in the underground and develop a resistant form like a cyst. With the aim to apply the chemigation for the control of ground-pearl, an assay was carried out in the laboratory. Tubes of 50 mm of PVC tubes (150 mm phi ) were cut and received a screen cover in one circular side. The tubes were then filled with natural soil, collected at Videira Experimental Station. Five tubes were put on the top of each other to form a test tube with distinct soil depths. Cysts of ground-pearl were put in the depths of 100 mm and 200 mm. The treatments were aldicarb 0.525 g i.a./plant (as a standard), metidathion 80 ml a.i./100 l, diazinon 90 ml a.i./100 l, imidacloprid 21 g a.i./100 l and destiled water (as an untreated control). The liquid insecticides and the water were applied as a surface drench at a rate of 20 l/m super(2). The best control was achieved with metidathion, resulting in 83.3% mortality after five months. Diazinon and imidacloprid resulted in 45.2% and 6.0% mortality respectively, while aldicarb resulted only in 1.2% mortality.

Hinkens, D.M., McElfresh, J.S. & Millar, J.G. 2001. Identification and synthesis of the sex pheromone of the vine mealybug, Planococcus ficus. Tetrahedron Letters 42 (2001): 1619-1621. [HinkenMcMi2001]

Notes: Sexually mature females of an important agricultural pest, the vine mealybug, Planococcus ficus, produces the monoterpene (S)-lavandulol and the corresponding ester, (S)-(+)-lavandulyi senecicate. The racemic ester was highly attractive to mature male mealybugs, whereas lavandulol was not. The naturally produced 2.5 blend of lavandulol and the ester also was no more attractive than the ester alone.

Hippe, C. & Frey, J.E. 2001 (1999). Biology of the horse chestnut scale, Pulvinaria regalis Canard (Hemiptera: Coccoidea: Coccidae), in Switzerland. Entomologica 33(1999): 305-309. [HippeFr2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] In 1997, many lime (Tilia spp.) and horse chestnut (Aesculus hippocastanum) trees in the centre of Zurich were found to be heavily infested by the horse chestnut scale, Pulvinaria regalis Canard. The biology of this introduced coccid was studied for one year. Crawlers hatched from the end of May and moved to the leaves of their host plants. There the nymphs settled and fed until Sept./Oct., when they migrated to adjacent twigs to overwinter as the 3rd-instar females. After the final nymphal moult in the spring, the adult female went through a period of rapid growth. Adult males appeared for a short period at the beginning of May but were rare. At this time, the females began to move to the main branches and the trunk of the tree, where they secreted a white ovisac consisting of wax filaments. Shortly after oviposition, the females died but remained attached to the ovisac. Two species of aphelinid (Coccophagus lycimnia (Walker) and, much less commonly, C. semicircularis (Förster)) emerged from parasitised scale nymphs in May (on twigs) and at the beginning of September (on leaves). The average rate of parasitisation of P. regalis was low (¾5%). No dipteran and only a few coccinellid predators were found during the sampling period.

Hodges, A.C. & Hodges, G. 2001. Notes on members of the Vespidae foraging on honeydew secretions from the European fruit lecanium, Parthenolecanium corni (Bouche). Journal of Entomological Science 36(3): 312-314. [HodgesHo2001]

Notes: Interactions among the insects found on P. corni honeydew are discussed. Ultracoelostoma spp. (Margarodidae) are also mentioned.

Hodgson, C. & Martin, J.H. 2001. Three noteworthy scale insects (Hemiptera : Coccoidea) from Hong Kong and Singapore, including Cribropulvinaria tailungensis, new genus and species (Coccidae), and the status of the cycad-feeding Aulacaspis yasumatsui (Diaspididae). Raffles Bulletin of Zoology 49(2): 227-250. [HodgsoMa2001]

Notes: Observations are presented on the current status of the cycad-feeding armoured scale, Aulacaspis yasumatsui Takagi (Diaspididae), in Singapore and Hong Kong. A new genus and species of soft scale, Cribropulvinaria tailungensis (Coccidae) from Aporusa dioica (Euphorbiaceae) is described from Hong Kong, with all stages except prepupa, and the possible relationships of this new genus discussed. The soft scale, Maacoccus cinnamomicolus (Takahashi), recently rediscovered in Singapore, has its adult female redescribed, 1st- and 3rd-instar nymphs described for the first time, and a lectotype designated.

Hosagoudar, V.B., Ganesan, R., Devi, D. & Ganesan, T. 2001. An observation on the sooty mould covered loquat tree. Journal of Economic and Taxonomic Botany 25(1): 13-14. [HosagoGaDe2001]

Notes: Erybotrya japonica is a fruit plant, introduced in hill stations of India, and is associated with scale insects. These fungal colonies prevent the entry of sunlight to the photosynthetic cells of the plant and also obstruct an immediate evaporation of the transpired water.

Houston, D.B. & Houston, D.R. 2000. Allozyme genetic diversity among Fagus grandifolia trees resistant or susceptible to beech bark disease in natural populations. Canadian Journal of Forest Research 30(5): 778-789. [HoustoHo2000]

Notes: American beech (Fagus grandifolia Ehrh.) trees resistant (R = 760) and susceptible (S = 681) to beech bark disease were located and mapped in nine natural stands in West Virginia (WV), Massachusetts (MA), Maine (ME), Nova Scotia (NS), and Prince Edward Island (PEI). Dormant bud tissue collected from each tree was examined by isozyme analysis to characterize the population genetic structure of R and S subpopulations mapped in each of four intensively sampled (R/S) stands, and five additional populations in which only R trees were sampled. Seventeen enzymes (with 9 polymorphic and 14 monomorphic loci) were analyzed to estimate variation across the spatial range of disease occurrence. All populations possessed significant levels of inter- and intra-population diversity; several parameters appeared to increase in magnitude from southwest to northeast across the range. Beech bark disease occurs after infestations of Cryptococcus fagisuga populations.

Hsu, J.C., Horng, S.B. & Wu, W.J. 2001. Spatial distribution and sampling of Aulacaspis yabunikkei (Homoptera: Diaspididae) in camphor trees. Plant Protection Bulletin (Taichung) 43(2): 69-81. [HsuHoWu2001]

Notes: Dispersion patterns of varied stages generated by Iwao's patchiness regression and Taylor's power law for Aulacaspis yabunikkei Kuwana in different sampling units of camphor trees, Cinnamomum camphora (L.), were compared. Taylor's power law provided a consistently good fit to the data, whereas the fit of Iwao's patchiness regression were erratic, and the values of aggregation index of Taylor's power law (1.76 to 2.65) were narrower than those of Iwao's (1.26 to 11.83), but both indices (b > 1.7 and beta > 1.2) indicate a clumped distribution pattern in all sampling units. Mean numbers of scales per leaf differed significantly (P < 0.05) between the lower and upper layer of the canopy, between old and young leaves, and between the underside and upper surface of leaves. The number of eggs corresponded closely to the total population, and this stage is best for precisely estimating the population. However, for the non-professional, the eggs are difficult to count. The number of female adult scale covers was easy to count and also corresponded very closely to the total population. Thus, female adult scale covers on the under leaf surfaces of old leaves were chosen as sampling targets to represent the scale population and for the decision making of pest management. Only scales with high density would damage camphor tree heavily, therefore, in fixed sampling plan, when there were 100 female scales on the undersides of leaves per twig in a tree, the optimum samples of 12 twigs per tree and 3 twigs per tree would reach 0.25 and 0.5 precision levels, respectively.

Hu, J.S., Sether, D., Melzer, M., Busto, J., Perez, E., Kislan, M., Dawson, W. & Karasev, A. 2000. Is mealybug wilt of pineapple a viral problem? Phytopathology 90(6) (Suppl.): S92. [HuSeMe2000]

Notes: Mealybug wilt of pineapple (MWP) is one of the most serious diseases in pineapple (Ananas sp.). We have produced specific monoclonal antibodies against two distinct pineapple mealybug wilt associated viruses (PMWaV). A RT-PCR assay was developed to detect and distinguish these two viruses. The genome of PMWaV 2 has been cloned and its sequence determined. The genome is typical of the monopartite closteroviruses. Sequence analysis shows that PMWaV 2 shares the highest nucleic acid homology with GLRaV-3. PMWaV 1 has also been cloned and six kilobases of its 5'-end region sequenced. The two PMWaVs are distinct from each other and share less than 50% nucleic acid homology based on available sequences. PMWaV 1 has highest sequence homology with GLRaV 1. Both PMWaVs can be acquired and transmitted by mealybugs (Dysmicoccus spp.). We have shown in transmission experiments that the presence of both PMWaV and mealybug exposure is necessary for the induction of MWP. The presence of the PMWaV in the absence of mealybug feeding does not induce MWP symptoms.

Hua, L.Z. 2000. (In Chinese). In: , List of Chinese Insects (Vol. 1). Zhongshan University Press, Guangzhou, China. 448 pp. [Hua2000]

Hunter, M.S. & Woolley, J.B. 2001. Evolution and behavioral ecology of heteronymous aphelinid parasitoids. Annual Review of Entomology 46: 251-290. [HunterWo2001]

Notes: In almost all species of parasitic wasps in the Coccophaginae, a sub-family of Aphelinidae, males have host relationships different from females. In these "heteronymous" species, females are generally endoparasitoids of sternorrhynchous Hemiptera, such as scale insects, mealybugs, and whiteflies. In contrast, males may be hyperparasitoids, developing in or on conspecific females or other primary parasitoids. In other species, females are endoparasitoids of whiteflies, and males are primary endoparasitoids of eggs of Lepidoptera. Males and females may both be primary parasitoids on the same species of scale insect hosts, but females develop as endoparasitoids, whereas males are ectoparasitoids. Here we review these life histories, focusing on examples of sexually dimorphic host relationships, development, and morphology. Coccophagine species may be sexual or parthenogenetic; we discuss reproductive modes and the interaction of sex ratio distorters with sex-specific host relationships. Sex allocation in the species in which males are hyperparasitoids involves choices of not what sex egg to lay, but whether to accept or reject a host of a given type; study in this area is reviewed as well as research in kin discrimination and ovicide. Last, we present the current understanding of phylogenetic relationships within this lineage and discuss hypotheses for the evolutionary origin of heteronomy in the Aphelinidae. Species mentioned include Chrysomphalus aonidum, Coccus hesperidum, Coccus proteae, Lecanium sp., Parasaissetia spp., and Quadraspidiotus ostreaeformis, Saissetia oleae.

Ismail, A.I. & Abdalla, E.F. 2001. Efficiency and residual activity of some selective compounds against Asterolecanium pustolans (pustulans?) Cock. and Ceroplastes rusci L. infesting fig trees. Annals of Agricultural Science (Cairo) 46(1): 355-364. [IsmailAb2001]

Notes: Field experiments were conducted in north Sinai, Egypt to evaluate some environmentally benign compounds (buprofezin, petroleum oils viz. Super Royal and kz oil, essential oils from Salvia sclarea, Ocimum basilicum and Mentha viridis (Mentha spicata)] and chlorpyrifos) against certain pests infesting fig orchards, Ficus carica. The treatments were able to arrest the development of nymphs and consequently, the emerged females were significantly suppressed. The insect growth regulator, buprofezin was the most effective compound against the pustule scale insect, A. pustulans followed by S. sclarea, Super Royal and kz oils, where the reduction in infestation reached 62.3, 49.2, 47.5 and 46.0%, respectively, throughout the whole 3 months of experiment. Adding 2.5% S. sclarea oil to the lower concentration of buprofezin (0.025%) displayed potentiation effect that destroyed the waxy scale, C. rusci population by about 87.5% and slightly exceeded the respective value for chlorpyrifos (84.3%). It should be noted that C. rusci population was more susceptible to all treatments either applied individually or in mixtures, than A. pustulans.

Jablonska, A. & Bilinski, S.M. 2001. Structure of ovarioles in adult queens and workers of the common wasp, Vespula germanica (Hymenoptera : Vespidae). Folia biologica (Krakow) 49(3-4): 191-198. [JablonBi2001]

Notes: The ovaries of the common wasp, Vespula germanica are polytrophic-meroistic and consist of 2-3 (workers) or 7 (queens) ovarioles. Scale insects are also mentioned.

Jalali, S.K., Singh, S.P. & Biswas, S.R. 2000. Parasitisation behaviour of Leptomastix dactylopii Howard (Hymenoptera: Encyrtidae) at various densities of Planococcus citri Risso. Journal of Entomological Research. New Delhi 24(2): 159-162. [JalaliSiBi2000]

Notes: The response of the encyrtid parasitoid, Leptomastix dactylopii Howard, to various densities of its host mealybug, Planococcus citri Risso, was investigated in the laboratory. Percent parasitization decreased, and adult parasitoid emergence increased, with the increase in host density. Development of immature stages did not differ among different density levels. The percentage of female progeny was greater at lower densities. A positive correlation between density and the number of adults emerged, and a negative correlation between density and parasitization was observed. A parasitoid:host ratio of 1:100 was optimal for parasitoid use and mass production of L. dactylopii.

Jansen, M.G.M. 2000. The species of Pulvinaria in the Netherlands (Hemiptera: Coccidae). (In English; Summary In Dutch). Entomologische Berichten (Amsterdam) 60: 1-11. [Jansen2000]

Notes: The occurrence of the species of the soft scale genus Pulvinaria in The Netherlands is discussed. Four species occur in the open, one is native (Pulvinaria betulae and three are introduced and established P. floccifera, P. hydrangeae and P. regalis. Two species are only known from interceptions during import inspections and from greenhouses (P. mesembryanthemi and P. psidii). A key to the species is given.

Jansen, M.G.M. 2001 (1999). An annotated list of the scale insects (Hemiptera: Coccoidea) of the Netherlands. Entomologica 33(1999): 197-206. [Jansen2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Sixty-six species of scale insects (Hemiptera: Coccoidea) have been recorded in The Netherlands up to August 1998. The species belong to 9 families, the most numerous of which are the Coccidae (19 species), the Pseudococcidae (15 species) and the Diaspididae (15 species). Nine species are recorded here for the first time: Matsucoccus matsumurae (Kuwana), Parthenolecanium rufulum (Cockerell), Phyllostroma myrtilli (Kaltenbach), Physokermes hemicryphus (Dalman), Quadraspidiotus marani Zahradnik, Quadraspidiotus perniciosus (Comstock), Steingelia gorodetskia Nassonov, Trionymus aberrans Goux and Trionymus tomlini Green. Carulaspis visci (Schrank) must be removed from the list.

Japoshvili, G.O. 2001 (1999). The parasitoid complex and population dynamics of the plum scale, Sphaerolecanium prunastri Fonscolombe, in Georgia. Entomologica 33(1999): 403-406. [Japosh2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The plum scale, Sphaerolecanium prunastri Fonscolomb, is found infesting stone fruits all over the Europe and in most of the countries of Asia and North America. Its population dynamics and parasitoid complex in Georgia are described and the latter compared with that in other countries in southern Europe. The present rates of parasitism are compared with those of an earlier survey and it is noted that there have been some changes, both in the composition and in effectiveness of the primary and secondary parasitoids. The main primary parasitoids are still Microterys hortulanus Erdös and Discodes coccophagus (Ratzburg) but, among the secondary parasitoids, Cerapterocerus mirabilis Westwood is now the most important species. It was found that the economic importance of S. prunastri had become reduced due to a reduction in the number of secondary parasitoids and an increase in numbers of primary parasitoids.

Jashenko, R.V. 2000. [Historical information on the employment of coccid species of the genus Porphyrophora Brandt (Coccinea, Margarodidae) in the ancient carmine industry and medicine, the possibility of their use in ethnographic research.] (In Russian). Tethys Entomological Research 2: 17-24. [Jashen2000]

Notes: A review of the historical data on the employment of Poryphyrophora species in carmine industry and medicine is given.

Jeyarani, S., Sridharan, S. & Sadakathulla, S. 2000. Effect of steam distillate extracts of resistant rice varieties on rice mealybug. Insect Environment 6(1) 14-15. [JeyaraSrSa2000]

Notes: Solutions of the steam distillates of mealybug [? Brevennia rehi] resistant rice cultivars (IR36, IR64, IR72 and Ptb21) were sprayed on the single tiller of the susceptible rice cultivar TN1 at 2000 ppm. Each plant was infested with 5 gravid mealybug females. Significantly fewer eggs were laid on plants treated with the distillates than on control plants. Toxicity tests on first instar nymphs revealed that the extracts of all cultivars were toxic, causing 54 to 68% mortality.

Johnson, C., Agosti, D., Delabie, J.H., Dumbert, K., Williams, D.J., Von Tschirnhaus, M. & Maschwitz, U. 2001. Acropyga and Azteca ants (Hymenoptera: Formicidae) with scale insect (Sternorrhyncha: Coccoidea): 20 million years of intimate symbiosis. American Museum Novitates (New York) No. 3335: 18 pp. [JohnsoAgDe2001]

Notes: Species of the genus Acropyga are rarely encountered subterranean ants that rely on mealybugs or aphids to provide their nutritional needs. Female Acropyga (Formicinae) alates of pantropical and Mediterranean species carry mealybugs with their mandibles while swarming and probably inoculate their new nests with these mealybugs. The natural history of Acropyga and other mealybug-tending ant species, a summary of the various reports of Acropyga and other mealybugs, and a new record from French Guiana are presented here. Also provided are a first report and description of Acropyga alates with mealybugs in Dominican amber dated to the Miocene, a discovery indicating that this intimate association and relatively uncommon behavior has existed for at least 15-20 million years. The mealybugs found with the Acropyga females in amber are related to the hypogaeic genera Eumyrmococcus Silvestri and Neochavesia Williams & Granara de Willink (Pseudococcidae, Rhizoecinae) and represent three new species of a new genus. The genus Electromyrmococcus and the species Electromyrmococcus abductus Williams, Electromyrmococcus inclusus Williams and Agosti, and Electromyrmococcus reginae Williams are described. A piece of Dominican amber containing workers of Azteca alpha Wilson (Dolichoderinae) and 23 scale insects is also presented and the significance of these specimens in Dominican amber is discussed.

Joubert, P.H., Daneel, M.S., Grove, T., & Pichakum, A. 2000. Progress towards integrated pest management (IPM) on mangoes in South Africa. Acta Horticulturae No. 509: 811-817. [JouberDaGr2000]

Notes: [Proceedings of the Sixth International Symposium on Mango, Pattaya City, Thailand, 6-9 April, 1999, Volume 2.] In South Africa, several insect pests are important in the production of mangoes. The mango scale, Aulacaspis tubercularis, mango seed weevil, Sternochetus mangiferae, and the fruit flies Ceratitis capitata, C. rosa and C. cosyra are all key pests and have to be controlled regularly. For the biocontrol of mango scale, an Aphytis sp. parasitoid and a predatory beetle, Cybocephalus binotatus, were imported from Thailand and established in several areas. Up to 46.3% parasitism of the mango scale occurred, while C. binotatus dispersed to several orchards surrounding the original release sites. Fenthion, which is used to control mango seed weevil, is extremely toxic to beneficial insects and alternative control measures were evaluated. Several methods e.g. systemic insecticides and egg parasitoids proved to be non-viable and finally a softer insecticide viz. endosulfan 475 g/kg SC, was evaluated for registration. For fruit fly control, most producers refrain from using full cover sprays and more IPM compatible bait sprays are applied. A monitoring system was developed for the important pests and the more judicious use of insecticides now facilitates IPM on mangoes in South Africa.

Joyce, A.L., Hoddle, M.S., Bellows, T.S. & Gonzalez, D. 2001. Oviposition behavior of Coccidoxenoides peregrinus, a parasitoid of Planococcus ficus. Entomologia Experimentalis et Applicata 98: 49-57. [JoyceHoBe2001]

Notes: The encyrtid parasitoid, Coccidoxenoides peregrinus has been used as a biological control agent against the mealybugs Planococcus citri and P. ficus. This study examined the behavior and host selection of C. peregrinus attacking P. ficus. Adult parasitoids were fed a 0.1% solution of acridine orange, a DNA binding dye used to label C. peregrinus eggs. In a choice test, adult parasitoids were offered equal numbers of first through fourth instars of P. ficus and behavior of C. peregrinus was filmed and analyzed. Acridine orange labeled ova of the parasitoids found within mealybug hosts fluoresced green under fluorescence microscopy and presence of fluorescing eggs in hosts was used to determine oviposition events. A time budget prepared for C. peregrinus indicated that this parasitoid spent the majority of its time searching (71.53%) and grooming (15.06%). The average probing duration over all instars which led to oviposition from single visits was 4.93% +/- 0.62 s. A total of 35.51% of probes from all attacks led to ovipositions, whereas 33.72% of single visits to hosts resulted in ovipositions. Detection of fluorescing acridine orange labeled eggs showed all instars of P. ficus were acceptable for oviposition by C. peregrinus. There was a significant preference to probe second, third, and fourth instars rather than first instars of P. ficus. Host feeding was not observed for this parasitoid.

Karaca, I., Senal, D., Colkesen, T. & Özgökçe, M.S. 2001 (1999). Observations on the oleander scale, Aspidiotus nerii Bouché (Hemiptera: Diaspididae) and its natural enemies on blueleaf wattle in Adana Province, Turkey. Entomologica 33(1999): 407-412. [KaracaSeCo2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The biology of Aspidiotus nerii Bouché and the overall efficiency of its natural enemies (the aphelinid parasitoid Aphytus melinus DeBach and the coccinellid predators Chilocorus bipustulatus (L.) and Rhyzobius lophantae (Blaisdell)) were studied. Forty leaves were collected at weekly intervals from 5 blueleaf wattle trees (Acacia saligna) from four compass bearings; all live and dead A. nerii and the number and stage of all parasitised scales were counted. There were two population peaks of A. nerii per year, in May/June and July/August. The number of parasitoids, however, fluctuated considerably, especially during the autumn and winter. The scale stage parasitised was primarily the adult female, followed by the pupae and then a few 2nd-instar nymphs. First-instar nymphs were never attacked by parasitoids but predators fed on all stages.

Karasev, A.V. 2000. Genetic diversity and evolution of closteroviruses. Phytopathology 90(6): S92. [Karase2000]

Notes: The family Closteroviridae comprises more than 30 plant viruses with flexuous, filamentous virions and includes representatives with either mono- or bipartite positive strand ssRNA genomes. Large genomes of closteroviruses, up to 19.3 kb, have apparent resemblance to genomes of arteri- and coronaviruses. Closteroviruses are transmitted semi-persistently by insects from three families of Homoptera, in infected plants are associated with phloem tissue, and demonstrate an astonishing genetic diversity which suggests an extensive, on-going evolution. Evolution of closteroviruses seems to proceed through duplications, acquisition of new genes, sometimes of host origin, and their subsequent diversification. Phylogenetic analyses of their replicative genes as well as the conserved HSP70 demonstrate that closteroviruses co-evolved with their insect vectors, resulting in three major lineages, i.e. aphid-, mealybug-, and whitefly-transmitted viruses. An amendment of the current classification will be discussed. Closteroviruses represent an ancient and diverse virus family which may pose threats to agriculture and needs serious attention.

Kaydan, M.B., Kozár, F., Yasar, B. & Erkiliç, L. 2001. Initial studies on Pseudococcidae fauna in Van Province of Turkey. Acta Phytopathologica et Entomologica Hungarica 36(3/4): 377-382. [KaydanKoYa2001]

Notes: Wild plants and cultivated crops from different habitats were surveyed and mealybug samples were collected from 1996 until 1998. Slides were prepared from these samples and nineteen species were identified, two of which had been previously identified. These species belonged to eight genera of the Pseudococcidae family. Antonina graminis, Atrococcus ingidens, A. saxatilis, Euripersia amnicola, Heliococcus bohemicus, H. saxatilis, Mirococcus inermis, Peliococcus bitubulatus, P. manifectus, P. terrestris, P. turanicus, P. unispinus, Phenacoccus affinis, P. karaberdi, P. tergrigorianae, Trionymus abberans and T. graminellus are recorded in Turkey for the first time.

Kim, K.W. 2000. [Non-chemical or low-chemical control measures against key insect pests and rats in the ginseng fields] (In Korean; Summary In English). Korean Journal of Applied Entomology 39(4): 281-286. [Kim2000]

Notes: Non-chemical or low-chemical control measures against key insect pests and rats in the ginseng field were studied from 1993 to 1999, in Suwon, Korea Republic. Application of chemicals in ginseng fields during oviposition had similar control effects on the Korean black chafer, Holotrichia diomphalia adults. Ginseng damage by the African mole cricket, Gryllotalpa africana adults were reduced considerably by application chemicals beside the ginseng field. The larvae of wheat wireworm, Ectinus sericeus, were attracted to potatoes in the ginseng field. The spread of the mealybug, Pseudococcus comstocki, was very slow in the ginseng field, indicating that it is possible to eradicate the early colonies effectively. The rat repeller, Dekur 500S showed a significant control effect of rats in the ginseng field.

Knight, A.L., Christianson, B.A. & Unruh, T.R. 2001. Impacts of seasonal kaolin particle films on apple pest management. Canadian Entomologist 133(3): 413-428. [KnightChUn2001]

Notes: The impact of multiple applications of the kaolin-based particle him M96-018 on the population density of selected pests of apple, Malus domestica (Borkh) (Rosaceae), and their natural enemy populations were measured in several Washington State orchards from 1997 to 1999. Fruit infestation by San Jose scale, Quadraspidiotus perniciosus (Comstock) (Hemiptera: Diaspididae), was significantly higher in treated plots than in untreated plots. No differences in pest densities occurred in plots treated for one versus two consecutive years.

Kosztarab, M. 2001 (1999). Status and future of human resources in coccidology. Entomologica 33(1999): 19-34. [Koszta2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] This paper reviews responses to a survey on the current status of researchers active in coccidology, with an emphasis on systematics but including work on morphology, biology, ecology, biological and chemical control of scale insects and on preparation of databases. The 39 responses provide a short assessment for 38 countries and 15 U.S. States, with the activities and/or whereabouts of about 243 researchers. Thirteen of the 32 active researchers (41%) will reach retirement age within five years; however, the average time to retirement for this group was 11.7 years. There has been a decline in the number of women in graduate training, while, from the 17 graduate students being trained in coccidology, only six are doing research in systematics.

Koteja, J. 2001. Essays on coccids (Hemiptera: Coccinea). Paleontology without fossils? (In English; Summary In Polish). Prace Muzeum Ziemi No. 46: 41-52. [Koteja2001]

Notes: Only a few phylogenetic conceptions that consider time were proposed for the scale insects, and they have been critically reviewed on the basis of the currently available fossils. Evident pre-Cretaceous coccid fossils are still lacking. Scale insects appeared "suddenly" as a large and diverse group in early cretaceous.

Koteja, J. 2001a (1999). Fossil coccids do exist. Entomologica 33: 35-36. [Koteja2001a]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Scale insects (Coccinea) have been discovered in almost all Tertiary and Cretaceous amber and in a few other deposits, and the number of known coccid fossils has increased from about 100 to approximately 900 in the last twenty years. This material represents a variety of forms and constitutes a reliable basis for serious palaeo-entomological studies.

Koteja, J. 2001b. Essays on coccids (Hemiptera: Coccinea): double function of Margarodid halteres. (In English; Summary In French). Annales de la Société Entomologique de France 27(3): 357-365. [Koteja2001b]

Notes: Direct observation of Porphyrophora polonica (L.) males showed that the margarodid halteres took part in reposing the wings, in addition to the role that they play in flight. This unusual behaviour is considered as an autapomorphie of the groundpearls which supports the view to treat them as a separate family, Margarodidae, distinct from other archeococcids.

Koteja, J. & Poinar, O. 2001. A new family, genus and species of scale insect (Hemiptera: Coccinea: Kukaspididae, new family) from Cretaceous Alaskan amber. Proceedings of the Entomological Society of Washington 103(2): 356-363. [KotejaPo2001]

Notes: A new genus and species of scale insect, Kukaspis usingeri is described from Cretaceous Alaskan amber and placed in a new extinct family, the Kukaspididae. This fossil is a derived member of the superfamily Orthezioidea, with six pairs of unicorneal eyes forming lateral rows, a scutum with a large subrectangular membrane, a tubular scutellum separated from the mesopostnotum by a large membrane, wings narrow with a clear posterior (claval) flexing line, but a reduced anterior one, narrow parallel-sided halteres; a unique waxy tail consisting of four soft filaments arising from the last abdominal tergite and a penial sheath divided into basal capsul and stylus with a hook-like apex. Relationships of this peculiar Lower Cretaceous form with both extant and extinct forms are discussed.

Kotikal, Y.K. & Kulkarni, K.A. 2000. Incidence of insect pests of turmeric (Curcuma longa L.) in northern Karnataka, India. Journal of Spices & Aromatic Crops 9(1): 51-54. [KotikaKu2000]

Notes: A roving survey conducted during 1996-97 at three phases of crop growth in turmeric (C. longa) in northern districts of Karnataka, India revealed the incidence of major insect pests such as scale insects (Aspidiotus curcumae) in all the areas.

Kozár, F. & Foldi, I. 2001. Description of a new species of Newsteadia from Corsica, France (Hemiptera, Coccoidea, Ortheziidae). (In English; Summary In French). Nouvelle Revue d'Entomologie 18(92): 157-160. [KozarFo2001]

Notes: Newsteadia susannae sp. n. is described from Corsica (France), increasing the number of the Newsteadia species known from the region to four. The new species was found together with N. floccosa (De Geer), which is widely distributed in the palaearctic region. N. susannae differs from N. floccosa in details of the antennal setae, by possessing numerous quadrilocular pores, and by lacking division between the wax plate bands on the mid-dorsum. The samples also contained N. floccosa, Ortheziola vejdovskyi and Arctorthezia cataphracta.

Kozár, F. & Konczné Benedicty, Z. 2001. Revision of Newsteadia (Homoptera: Coccoidea) of the Nearctic and Neotropic regions, with descriptions of new species. Acta Phytopathologica et Entomologica Hungarica 36(1-2): 123-142. [KozarKo2001]

Notes: Newsteadia borhidii new sp., N. brasiliensis new sp., N. costaricaensis new sp., N. floridensis new sp., N. minima, N. morrisoni new sp., N. trisegmentalis, N. tristani, and N. tropicalis new sp. are described and the new species are illustrated. Key to adult females of Newsteadia found in these regions is provided.

Kozár, F. & Konczné Benedicty, Z. 2001a. Ortheziola of Asia with the descriptions of three new species, and world distribution of the genus (Homoptera : Coccoidea, Ortheziidae). Acta Zoologica Academiae Scientiarum Hungaricae 47(1): 15-25. [KozarKo2001a]

Notes: Three new Ortheziola species (O. peregovitsi, O. vietnamiensis, O. matskasii) are described from Asia. By these descriptions the number of the known Ortheziola species of the World increases to 23. Distribution records and zoogeographical considerations are given. The new species represent a link between the Palaearctic, Oriental and Ethiopian Regions. The records gave a new insight into the species richness of this genus in the World.

Kozár, F. & Miller, D.R. 2000. World revision of Ortheziola Šulc (Homoptera: Coccoidea: Ortheziidae) with descriptions of eleven new species. Systematic Entomology 25: 15-45. [KozarMi2000]

Notes: Soil samples and museum collections were analysed from all zoogeographic regions of the world. From this material eleven new species of Ortheziola are described from Africa and the U.K., five additional species are redescribed and a key is presented for the identification of all sixteen species. A slightly modified concept of the genus is presented and several new characters are given as diagnostic of the genus. A phylogenetic hypothesis is given based on analysis of the morphological features of adult females. Results show that Ortheziola is most diverse in eastern Africa where relatively more advanced species occur. Basal species occur primarily in western Africa and Asia.

Kozár, F. & Miller, D.R. 2001 (1999). Observations on collecting scale insects (Hemiptera: Coccoidea). Entomologica 33(1999): 243-250. [KozarMi2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Scale insects have been primarily collected visually. Because scales are usually firmly attached to the host substrate, most mass-collecting techniques are ineffective. This paper provides information on the use of Berlese funnels, DVAC suction, pit-fall traps, sweeping, beating and screening for collecting scale insects and provides observations on how these methods compare with visual collecting methods.

Kozár, F. & Seprös, I. 2001. New scale insect pests (Homoptera: Coccoidea: Coccidae) on urban ornamentals. (In Hungarian; Summary In English). Növényvédelem 37(9): 441-444. [KozarSe2001]

Notes: Two important coccid species were found during this survey on urban ornamental plants: camellia scale (Chloropulvinaria floccifera) and hydrangea scale (Eulpulvinaria hydrangea). The latter species is new for the Hungarian fauna and both species are new on the list of the major scale insect pests. These species may attack various shrubs and trees, such as Euonymus, Taxus, Celtis, Sophora, Aesculus, Acer, Tilia and other ornamentals. Close monitoring of their distribution and population increases are recommended. Pulvinaria regalis is another species likely to occur.

Krasny, M.E. & di Gregorio, L.M. 2001. Gap dynamics in Allegheny northern hardwood forests in the presence of beech bark disease and gypsy moth disturbances. Forest Ecology and Management 144(1-3): 265-274. [KrasnyDi2001]

Notes: To determine changes in gap dynamics over a 6-year period in forests affected by beech bark disease (a disease complex of the scale insect, Cryptococcus fagisuga, and the fungi, Nectria) and gypsy moth infestations line transect surveys of gaps and gap makers were conducted in three Allegheny hardwood forests in central New York, USA. Gap makers were defined as any trees whose death or loss of > 50% of the branches caused an opening in the canopy, and were divided into three categories: decline (standing live trees with > 50% branch loss), standing dead, and treefall. Total land area in gaps increased significantly (P_0.00) from 19.7 to 31.9% during the 6-year study period. American beech (Fagus grandifolia) represented 52.4% of the gap makers in 1990 and 1996. Of the total beech gap makers, 30% were decline and 53% were treefall gap makers in 1990, but by 1996, only 8% were decline as compared to 76% treefall gap makers. In contrast, the majority of oak (Quercus spp.) gap makers were standing dead in 1990 and 1996. Although the rate of gap formation slowed at the end of this study, the predominance of beech in the subcanopy and regeneration layers pose persistent challenges for forest managers.

Kreiter, P., Dijoux, L., Donnadieu, F., Malausa, J.C. & Quilici, S. 2000. [Tentative introduction of Arrhenophagus chionaspidis (Hymenoptera, Encyrtidae) into the Alpes-Maritimes for control of the white peach scale.] (In French; Summary In English). Bulletin Mensuel de la Société Linnéenne de Lyon 69(6): 121-126. [KreiteDiDo2000]

Notes: [Original title: Tentative d'introduction dans les Alpes-Maritimes de la souche réunionnaise d'Arrhenophagus chionaspidis (Hymenoptera, Encyrtidae) pour lutter contre la Cochenille blanche du mûrier.] This parasitoid on Pseudaulacaspis pentagona has been introduced from Reunion Island to the Alpes-Maritimes. Biological characteristics are discussed.

Kubiriba, J., Legg, J.P., Tushemereirwe, W. & Adipala, E. 2001. Disease spread patterns of Banana streak virus in farmers' fields in Uganda. Annals of Applied Biology 139(1): 31-36. [KubiriLeTu2001]

Notes: Three surveys were conducted to establish the disease spread patterns of Banana streak virus (BSV) in farmers' fields in Uganda. Transects were traced both across the fields and from infection foci within a field. BSV incidence in adjacent quadrants was also determined to quantify statistically the spatial relationships of infected plants in the fields. Severity assessment along transects across fields revealed clusters of plants with moderate to high severity and clusters of plants with no BSV or low severity. Symptom severity decreased away from foci of infection (b =-0.014; P= 0.0081). Observed frequency of infected quadrant counts differed from corresponding expected frequency of infected quadrant counts (Poisson's distribution, chi (2); p less than or equal to 0.01). BSV-infected plants, therefore, were aggregated in well-established fields. Aggregation of infected plants in farmers' fields and the decrease of severity away from infection foci suggest the likely involvement of a slow moving vector (mealybugs) in BSV transmission.

Kubiriba, J., Legg, J.P., Tushemereirwe, W. & Adipala, E. 2001a. Vector transmission of Banana streak virus in the screenhouse in Uganda. Annals of Applied Biology 139 (1): 37-43. [KubiriLeTu2001a]

Notes: Although mealybug transmission of Banana streak virus (BSV) by Planococcus citri and Saccharicoccus sacchari has been demonstrated elsewhere, these mealybugs have not been identified on bananas in Uganda and their role and that of other agents in BSV transmission is not well documented. Insect samples were collected from banana farms in sites with low, moderate and high BSV infections in Uganda. Subsequently, live mealybugs and aphids were again collected and used in acquisition, retention and transmission tests, and BSV diagnosed using TAS-ELISA. Dysmicoccus brevipes (pineapple mealybug), S. sacchari (sugarcane mealybug ) and Pentalonia nigronervosa (banana aphid) were the most abundant insect species from banana fields sampled. Abundance of D. brevipes was positively and significantly correlated with BSV incidence unlike that of P. nigronervosa. Transmission studies in the screenhouse showed that mealybugs acquired BSV one day after feeding on virus sources and approached optimum acquisition after the third day. Pineapple and sugarcane mealybugs retained BSV up to 5 days from the day of transfer from the virus source. BSV was first detected in the recipient banana plants 4 wk after transmission using pineapple mealybug and 6 wk after inoculation using sugarcane mealybug. Under screenhouse conditions, both mealybugs therefore appear to transmit BSV semipersistently.

Kunca, A., Zubrik, M. & Leontovyc, R. 2000. Contribution to the knowledge of beech bark disease in the Magura working-plan area, West Carpathian mountains. (In English; Summary In Slovakian). Lesnicky Casopis 46(1):59-67. [KuncaZuLe2000]

Notes: Occurrence of beech bark necrosis, the beech scale insect (Cryptococcus fagi [Cryptococcus fagisuga]) and feeding by larvae of the butterfly Ectoedemia liebwerdella, and several site attributes, were evaluated for 1163 beech (Fagus sylvatica) trees in a 1800 ha area at Strazovskych in the West Carpathian region of Slovakia. The influence of the various factors measured on the occurrence of beech bark necrosis was analysed. Neither the beech scale insect nor feeding by E. liebwerdella larvae had a significant influence on the presence of bark necrosis. Unfavourable site conditions, such as shallow soils, steep slopes and south to east exposures had some influence on the occurrence of the disease.

Lagowska, B. 2000. Puto superbus (Leonardi, 1907) (Homoptera: Pseudococcidae) new to the Polish fauna. Polskie Pismo Entomologiczne 69: 3-6. [Lagows2000]

Notes: Puto superbus (Leonardi) is recorded for the first time from Poland. It represents the second indigenous species to Poland in the genus. Data on distribution, host plants and life history are given.

Lagowska, B. 2000a. [The scale insects (Homoptera, Coccinea) of ornamental coniferous plants in Poland: pest status and control.] Proceedings of the XVth Czech and Slovak Plant Protection Conference 379-380. [Lagows2000a]

Notes: [Conference held in Brno, September 12-14, 2000.] Nine species belonging to three families are listed. Four species are economically important: Parthenolecanium fletcheri and P. pomeranicum (Coccidae), Carulaspis juniperi (Diaspididae) and Planococcus vovae (Pseudococcidae). Plant hosts, biological and life history notes, and suggestions for chemical control are included.

Lagowska, B. 2001. Morphological variation in adult female Coccus hesperidum L. (Hemiptera: Coccoidea: Coccidae) in Poland. Entomologica 33: 105-112. [Lagows2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] This paper compares the variation of ten morphological characters of adult female C. hesperidum L. collected off (i) five different host plants and (ii) the stems, petioles and both leaf surfaces of Citrus, all from glasshouses in Poland. Four measured and six meristic characters were selected for biometric study. The mean and range of each of these characters varied between host plant species and with their position on Citrus. The main characters which varied between host species were the size and shape of the dorsal setae and the distribution and frequency of dorsal and ventral tubular ducts. Specimens off different parts of Citrus varied mainly in the frequency of the dorsal tubular ducts and the spiracular disc-pores. Pocket-like sclerotisations were noted for the first time in C. hesperidum; when present, they were in the stigmatic areas, close to a dorsal tubercle.

Lagowska, B. & Gawlowska, J. 2001. [Scale insect damage on Thuja and yew trees.] (In Polish; Summary In English). Ochrona Roslin 45: 1-2, 36-36. [LagowsGa2001]

Notes: [Original title: Szkodliwosc misecznikow na tujach i cisach.] Three species of scale insect attacking ornamental trees and shrubs in Poland, are described: Parthenolecanium fletcheri (occurring most frequently on Thuja and yew), P. corni and P. pomeranicum (common on yew). The life cycle of each pest and the most effective methods of control are described.

Lambdin, P.L. 2001. Morphology of the second instars of Polea martini and Polea selangorae (Coccoidea: Asterolecaniidae). Entomological News 112(3): 167-172. [Lambdi2001]

Notes: The second instars of the pit scales (Asterolecaniidae) Polea martini and P. selangorae are described and illustrated with measurements included for the various morphological structures. Both species were collected on species of Calophyllum sp. from Borneo and Singapore, respectively. These species are morphologically similar to immatures of Grammococcus in the New World. Polea martini is distinguished from P. selangorae by the presence of an anal ring with 6 setae, more multilocular pores in the dorsal pore clusters, and differences in the tubular ducts.

Lambdin, P.L. & Joshi, P.A. 2001. Haemocytes of three scale insect species: Phenacoccus gossypii Townsend & Cockerell, Pseudococcus longispinus (Targioni Tozzetti) and Dactylopius confusus (Cockerell) (Hemiptera: Coccoidea). Entomologica 33: 113-117. [LambdiJo2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] An evaluation of the haemocytes in the cochineal scale, Dactylopius confusus (Cockerell), was completed and compared with those found in the mealybugs Phenacoccus gossypii Townsend & Cockerell and Pseudococcus longispinus (Targioni Tozzetti) to assess the potential sites of the dye pigment source. Four basic cell types were found in the two pseudococcids and five in the cochineal scale. The cell types common to all species included: prohaemocytes, oenocytoids, typical granulocytes and plasmatocytes. In addition, a modified granulocyte (poly-glyco-based granulocyte) was found to be specific to the cochineal scale and this produced rough endoplasmic reticulum derived granules that may be the source for the synthesis of carminic acid.

Lambdin, P.L. & Podsiadlo, E. 2001. Descriptions of the first instars of Lecanodiaspis brookesae and L. crassispina (Coccoidea: Lecanodiaspidae). Entomological News 112(5): 287-293. [LambdiPo2001]

Notes: The first instars of Lecanodiaspis brookesae and L. crassispina from Australia are described and illustrated. Lecanodiaspis brookesae is distinguished by the presence of fleshy anterior and posterior spiracular setae, ten labial setae, and two setae on the anal plates, while L. crassispina is differentiated from other species of Lecanodiaspis by fleshy anterior and spine-like posterior spiracular setae, ten labial setae, and three setae on the anal plates. A modified key is provided to separate these species from other known species in this taxon.

Latta, S.C., Gamper, H.A. & Tietz, J.R. 2001. Revising the convergence hypothesis of avian use of honeydew: evidence from Dominican subtropical dry forest. Oikos 93(2): 250-259. [LattaGaTi2001]

Notes: A honeydew-producing scale insect (family Margarodidae, tribe Xylococcini, genus Stigmacoccus) was found associated with the tree Bursera simaruba in subtropical dry forests at elevations of 100-400 m on the south side of the Sierra de Bahoruco, Pedernales Province, Dominican Republic. At two study sites, 91% of Burseia trees supported locally dense populations of Margarodidae. Fifteen species of birds were observed foraging on the honeydew, but most observations were of the winter resident Cape May warbler (Dendroica tigrina) and black-throated blue warbler (D. caerulescens), and the permanent resident bananaquit (Coereba flaveola) and black-crowned palm tanager (Phaenicophilus palmarum). The Cape May warbler actively defended the honeydew resource but frequency of use of honeydew was influenced by the close presence of flowering agave and scale insect density. Data suggest that honeydew may be a critical component of the diet of this species especially during the late winter dry-season. Hymenopteran insects also were observed feeding on honeydew, but rates of consumption did not approach that of avian species. The occurrence of this phenomenon in Dominican dry forest is discussed in light of the convergence hypothesis of bird use and defense of homopteran honeydew in which it is proposed that birds are able to maintain relationships with scale insects in moist, warm temperate forests because it is in these climates where ant abundance is low. We suggest that our observation of a well-developed bird-homopteran system in classic subtropical dry forest supports the proposed mechanism of reduced competition with ants allowing bird use of honeydew, but we suggest that a broader array of especially insular habitats which may be relatively depauperate in terms of ants also can be expected to support bird-homopteran systems.

Le Ru, B. & Makosso, J.P.M. 2001. Prey habitat location by the cassava mealybug predator Exochomus flaviventris: Olfactory responses to odor of plant, mealybug, plant-mealybug complex, and plant-mealybug-natural enemy complex. Journal of Insect Behavior 14(5): 557-572. [LeRuMa2001]

Notes: Exochomus flaviventris Mader is considered to be the most active predator of the cassava mealybug Phenacoccus manihoti Matile-Ferrero in Central Africa. The response of experienced gravid female coccinellids to the odor of cassava plant (var. Zanaga), unparasitized mealybugs, plant-mealybug complex with or without feeding prey (parasitized or not), and plant-mealybug complex with or without conspecific coccinellids was investigated in a Y-tube olfactometer. The odor of uninfested cassava plants was not more attractive than clean air. Dual-choice tests revealed that mealybug-infested plants were preferred to mealybugs alone and mealybug-damaged plants and were the major sources of volatiles that attract female coccinellids to the microhabitat of its prey. The emission of volatile chemicals did not appear to be limited to the infested parts of the plant but did occur systemically throughout the plant. The presence of conspecific coccinellid larvae or adult males did not modify the attractiveness of the mealybug-infested plants. However, when an infested plant with conspecific predator females (alone or with conspecific males) was compared to an infested plant or infested plant with conspecific males, E. flaviventris females showed a preference for the last two sources of odor. The infested plant with conspecific males was also preferred to the uninfested plant with conspecific females. In addition, the odor of conspecific males was preferred to that of conspecific females. Female predators preferred the plant infested with unparasitized mealybugs to the plant infested with mealybugs previously parasitized. These results showed that E. flaviventris females use herbivore-induced plant volatiles during foraging and can detect via olfaction the presence of conspecific gravid females and parasitized prey, thus assessing patch suitability from a distance.

Lima, I.M.M. & Gama, N.S. 2001. [Record of host plants (Cactaceae) and new dissemination strategy of Diaspis echinocacti (Bouche) (Hemiptera: Diaspididae), Prickly-Pear-Scale, in the states of Pernambuco and Alagoas, Brazil] (In Portuguese). Neotropical Entomology 30(3): 479-481. [LimaGa2001]

Notes: [Original title: Registro de Plantas Hospedeiras (Cactaceae) e de Nova Forma de Disseminaçao de Diaspis echinocacti (Bouche) (Hemiptera: Diaspididae), Cochonilha-da-Palma-Forrageira, nos Estados de Pernambuco e Alagoas.] Opuntia ficus-indica and Nopalea cochenillifera (Cactaceae) are alternative food sources for cattle in drought areas of the Northeastern Region of Brazil. Diaspis echinocacti (Bouche) is the unique pest of these Cactacea species and it was previously detected on the following native host plants: Cereus jamacaru, Melocactus sp., Opuntia dillenii and O. inamoena. The new record of native hosts are Cereus gounellei, C. squamosus and the exotic C. hildemannianus from drought areas; Pereskia grandifolia and Epiphyllum oxypetalum, native species from tropical forest used as ornamental plants. D. echinocacti has not infested O. monacantha in the field and in laboratory bioassays. The dissemination of the neonate scale insect transported on the body surface of the predators Chilocorus nigrita and Zagreus bimaculosus Coccinellidae: Chilocorinae) was recorded for the first time.

Lin, T., Liu, K.Y., Liu J.X., Xiao, F., Deng, L.W., Yan, S.C., Li, F.R. & Li, C.D. 2000. [The population dynamics and prediction technique of Lepidosaphes salicina.] (In Chinese; Summary In English). Journal of Northeast Forestry University 28(6): 52-54. [LinLiLi2000]

Notes: A periodic systematic survey of the death rate of the first instar settled nymphs, pupae and adult females as well as the development progress of the newly hatched nymphs, second instar nymphs, prepupae and pupae was performed in Hongqi Forest Farm of Daqing City, Heilongjiang Province.

Lit, I.L. & Gullan, P.J. 2001. Comparative morphology of the anal tubercle and associated structures of some lac insects (Hemiptera: Coccoidea: Kerriidae). Entomologica 33: 119-126. [LitGu2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Specimens of Austrotachardia acaciae (Maskell), Austrotachardia sp. ex Cassinia spp., Kerria lacca (Kerr) and Paratachardina decorella (Maskell) were examined under the scanning electron microscope (SEM). SEM images were compared with observations under the light microscope. The anal tubercles and associated structures of each species are described and compared. The presence of what appears to be vestiges of an anal cleft in Paratachardina is interpreted as suggesting the possible evolution of the lac insects from a lecanoid ancestor that possessed an anal cleft. The results provide an easier interpretation of these characters under the light microscope.

Lit, I.L., Caasi-Lit, M. & Talidong, C.V. 2001 (1999). The production of long anal filaments by the bamboo node mealybug, Antonina sp. (Hemiptera: Coccoidea: Pseudococcidae), as a response to lack of attending ants. Entomologica 33(1999): 311-316. [LitCaTa2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Bamboo node mealybugs, Antonina spp., in the absence of attending ants, were observed to produce long waxy filaments both in the field and in the greenhouse. In contrast, ant-attended mealybugs had only very short filaments or none at all. Ant exclusion experiments confirmed the field observations. The available data suggest that the long filaments are an adaptation for the dispersal of honeydew in the absence of solicitous ants, so as to avoid drowning in the accumulating honeydew or suffocation due to development of sooty moulds.

Liu, J.X., Liu, K.Y., Lin, T., Wang, H.Y. & Liu, D.Y. 2000. [Regulations of the outbreak of Lepidosaphes salicina.] (In Chinese; Summary In English). Journal of Northeast Forestry University 28(6): 48-51. [LiuLiLi2000]

Notes: Distribution of L. salicina on their hosts is discussed. The death rate was high in hot summers and severe winters. Investigation of the relationship between the occurring area of oyster scale and meteorological factors such as humidity and temperature in ten cities and counties in Heilongjiang province formed a multiple linear regression model for the prediction of the occurring area of oyster scale. Acarids, ladybeetles, parasitic wasps and parasitic flies were found to be natural enemies.

Liu, S.Q. & Xu, H.Z. 2000. [Integrated control and characteristics of the main poplar diseases and insect outbreaks in Siyang County.] (In Chinese; Summary In English). Journal of Jiangsu Forestry Science and Technology 27(6): 33-35. [LiuXu2000]

Notes: The characteristics and integrated control are briefly discussed of the 2 fungal diseases and 5 insect pests of poplars [Populus] found in Siyang County, Jiangsu Province, China. Among the insect pests is Drosicha corpulenta.

Lizaso, M.T., Moneo, I., García, B.E., Acero, S., Quirce, S. & Tabar, A.I. 2000. Identification of allergens involved in occupational asthma due to carmine dye. Annals of Allergy, Asthma & Immunology 84(5): 549-552. [LizasoMoGa2000]

Notes: Carmine has been implicated as an etiologic agent of occupational asthma, but the allergens involved have not been yet identified. An in vitro study based in SDS-PAGE and IgE immunoblotting with carmine and cochineal extracts was performed. Sera from three carmine dye workers diagnosed with occupational asthma induced by carmine dye and from seven nonatopic subjects were used. Three proteins of around 30, 28, and 17 kD in raw cochineal extract and another protein of 50 kD in the boiled one were demonstrated by SDS-PAGE. Two proteins of around 50 and 28 kD were observed in the carmine extract by the same technique. Specific IgE binding bands at 17 kD in cochineal raw extract, at 50 kD in the boiled one, and at 28 kD in carmine extract were demonstrated by IgE immunoblotting. We have identified three allergens of around 17, 28, and 50 kD implicated in occupational asthma of three carmine workers.

Lo, P.L. & Chapman, R.B. 2001. Predation by Halmus chalybeus (Coleoptera: Coccinellidae) on Ceroplastes destructor and C-sinensis (Hemiptera: Coccidae: Ceroplastinae) infesting citrus in Northland, New Zealand. Biocontrol Science and Technology 11(1): 57-66. [LoCh2001]

Notes: Predation by Halmus chalybeus (steelblue ladybird) on two species of wax scale was studied on citrus orchards in Northland, New Zealand. Field experiments using an exclusion technique of enclosing citrus branches in bags, found that larval and adult H. chalybeus preyed on first and second instars of both Ceroplastes destructor (white wax scale) and C. sinensis (Chinese wax scale), but not third instar C. destructor. Third instar C. sinensis and adults of both species were not tested but are rarely, if ever, attacked by H. chalybeus. The sampling of scale and ladybird populations and Weld experiments showed that few scales of either species survived past the second instar stage where H. chalybeus was numerous. The feeding rate of H. chalybeus on settled first and second instar scales was estimated inside bagged branches. Adults consumed on average 15.6 C. destructor and 13.3 C. sinensis per day per ladybird, while larvae ate 9.7 C. destructor per day. These feeding rates accounted for the experimental reduction of scale populations. H. chalybeus is a useful natural enemy for the control of C. destructor and C. sinensis when its activity is not disrupted by pesticides.

Longo, S., Marotta, S., Pellizzari, G., Russo, A. & Tranfaglia, A. 2001. A zoogeographical analysis of the Italian scale insect fauna. Entomologica 33: 229-230. [LongoMaPe2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. The results of a zoogeographical analysis of the Italian scale insect fauna, which currently includes 365 species were presented. Eleven species were included in the analysis, either because they were poorly described and were therefore of doubtful identity or because they had not been recorded since their original description. This analysis divided the Italian scale insect fauna into three major groups according to the distributional patterns currently used in faunistic studies in the Western Palaearctic Region: A) Species primarily of the Western-Palaearctic Region (226 species). This group included the following sub-groups: i) species widely distributed in the Holarctic region (115 spp.); ii) species widely distributed in Europe (52 spp.); iii) species widely distributed in the Mediterranean Basin (57 spp.), and iv) Afrotropical or Oriental species also present in the Mediterranean area (2 spp.). B) Cosmopolitan species or cultural immigrants (111 spp.). C) Endemic species (17 spp). Each major group or sub-group included several species which had a more restricted distribution pattern. The analysis revealed that the Italian scale insect fauna (with the exclusion of the cosmopolitan species, which are of little zoogeographical interest), was represented mainly by widely distributed species in the Holarctic region (31.5% of total scale fauna); those which were widely distributed in the Mediterranean Basin (15.6%) and those widely distributed in Europe (14.25%). The Afrotropical or Oriental species represented only 0.5% of the total. A brief comment on each group was given.

Lorenzo Fernández, J.M., Prendes Ayala, C. & Lorenzo Bethencourt, C.D. 2001. [Following of populational dynamic of Dysmicoccus grassii (Leonardi) in Musa acuminata Colla. Subgroup Cavendish cv. Petit naine.] (In Spanish). Boletín de Sanidad Vegetal, Plagas 27(1): 85-101. [LorenzPrLo2001]

Notes: [Original title: Seguimiento de la dinámica poblacional de Dysmicoccus grassi (Leonardi) (Homoptera: Pseudococcidae) en Musa acuminata Colla, subgrupo cavendish cv. Pequeña enana.] The optimum number of plants was determined for estimating the extent of D. grassii infestation. Maximum and minimum temperatures and relative humidity discussed in relation to the persistence and recurrence of this pest.

Lotfalizadeh, H. & Ahmadi, A.A. 2000. Natural enemies of cypress tree mealybug, Planococcus vovae (Nasonov), and their parasitoids in Shiraz, Iran. (In English; Summary In Persian). Iran Agricultural Research 19(2): 145-154. [LotfalAh2000]

Notes: In a survey conducted in 1998, 15 species of natural enemies attacking cypress tree mealybug, Planococcus vovae were found in Shiraz, Iran. These included two parasitoids, Anagyrus pseudococci and Dusmetia fascipennis. The most common predators included Exochomus spp., Hyperaspis polita, Nephus bipunctatus, Chrysoperla carnea, Suarius fedtschenkoi, Dicrodiplosis manihoti Harris and Geocoris quercicola. The effectiveness of these natural enemies was reduced by the hyperparasitoids, Marietta picta and Chartocerus spp., by the parasite Homalotylus ephippium attacking coccinellids, and by the parasites Cheiloneurus ceroplastis, Ooencyrtus kuvanae, Isodromus atriventris, I. aff. vinulus, Catolaccus crassiceps and Pachyneuron concolor [P. muscarum] attacking chrysopids.

M'Hamed, T.B. & Chemseddine, M. 2001. Assessment of temperature effects on the development and fecundity of Pullus mediterraneus (Col., Coccinellidae) and consumption of Saissetia oleae eggs (Hom., Coccoidea). Journal of Applied Entomology 125(9-10): 527-531. [MHamedCh2001]

Notes: Eggs, larval and nymphal periods and fecundity of Pullus mediterraneus were examined under 16 h light : 8 h dark combined with six constant temperatures: 15, 20, 25, 30, 35 and 40 degrees C. Eggs of Saissetia oleae were used as prey. The developmental time at 15, 20, 25, 30 and 35 degrees C was 17.23, 4.5, 2.64, 1.67, 1.28 days for eggs and 98.47, 68.88, 53.94, 28.96, 36.51 days for larval-pupal duration, respectively. At 7 degrees C no eggs hatched, and at 40 degrees C all the stages died after 36 h of maximum exposure except the three last stages. The fecundity of females rearing at different temperatures ranged between 1.7 eggs at 15 degrees C and 601.86 eggs at 30 degrees C. The pre-oviposition period ranged between 23.75 days at 15 degrees C and 3.47 days at 35 degrees C. The consumption of S. oleae eggs by the larvae reached 597.69 eggs during the pre-imaginal development. Females attacked more eggs than males averaging 77.69 +/- 22.34 eggs per 4 day period compared with 46.97 +/- 10.12 eggs per 4 day period for males.

Mahmoud, F.A., Hamdy, M.K. & Hegazi, A.G. 2001 (1999). Antimicrobial activity of secretory materials of some scale insects. Entomologica 33(1999): 441. [MahmouHaHe2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. The antimicrobial activity of the secretory materials associated with the test or wax covers of five scale insects was tested for Gram-positive and Gram-negative bacteria. The scale insects studied were the margarodid Icerya aegyptiaca (Douglas), the soft wax scale Ceroplastes rusci (Linnaeus) and three hard scales: Aonidiella aurantii (Maskell), Lepidosaphes beckii Nerwman and Parlatoria zyziphi (Lucas). The Gram-positive bacteria found included Staphylococcus aureus and Streptococcus agalactia, while the Gram-negative bacteria included Escherichia coli, Pseudomonas aeruginosa and Corynebacterium pseudotuberculosis. The results of these observations revealed that there was some highly significant antimicrobial activity associated the secretary materials of the diaspidids A. aurantii, L. beckii and P. zyziphus. The antimicrobial activity observed varied according to type of the pathogenic bacteria it was used against. Thus, the activity from all three species was effective against Staphylococcus aureus, but that of A. aurantii and P. zyziphus was less effective against Corynebacterium pseudotuberculosis.

Malsch, A.K.F., Kaufmann, E., Heckroth, H.-P., Williams, D.J., Maryati, M. & Maschwitz, U. 2001. Continuous transfer of subterranean mealybugs (Hemiptera, Pseudococcidae) by Pseudolasius spp. (Hymenoptera, Formicidae) during colony fission? Insectes Sociaux 48: 333-341. [MalschKaHe2001]

Notes: In west- and east-Malaysian lowland rainforests, three Pseudolasius spp. are found in trophobiotic relationship with a total of eight scale insect species in six genera feeding at the roots of plants. Intimate associations going beyond pure trophobiosis could be discovered with five species of root mealybugs: Planococcoides sp., Maconellicoccus multipori and three species of Rhizoecus. All three Pseudolasius species carry their pseudococcids when disturbed, during nest movements and to feeding sites. Moreover, the Pseudolasius species discussed permanently keep adults and immature instars of their mealybug partners within their nests.

Malumphy, C. 2001. Scale insects (Hemiptera, Coccoidea) intercepted on imported plant material and recent introductions in England and Wales. Entomologica 33: 230. [Malump2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. There have been almost 300 species of Coccoidea intercepted on imported plants and plant produce in England and Wales since 1968, which is more species of Coccoidea intercepted than any other superfamily of invertebrates. For example, in 1995, a third of all species intercepted on imported plant material were Coccoidea. This makes scale insects the most significant superfamily of invertebrates being dispersed in the international plant trade, in terms of numbers of species. The most frequently intercepted species are listed and the reasons why they are so commonly transported briefly discussed. Despite the large number of exotic scale insects entering Britain very few have become established and even fewer have become widespread. New introductions of exotic species reported in non-commercial premises in Britain since 1968 are also listed. Finally, recent outbreaks of exotic pest species at commercial nurseries are given, all of which have been successfully eradicated.

Mani, M. & Krishnamoorthy, A. 2000. Predatory Potential and Development of the Australian Ladybird Bettle, Cryptolaemus montrouzieri Muls. on the Spiralling Whitefly, Aleurodicus dispersus Russel. Entomon 24(2): 173-176. [ManiKr2000]

Notes: The Australian ladybird beetle Cryptolaemus montrouzieri Muls. was found commonly preying on the spiralling whitefly, Aleurodicus dispersus Russel in South India. Predatory potential and development of the larvae of C. montrouzieri was studied on the nymphs of the spiralling whitefly. The duration of first, second, third and fourth instar larva of C. montrouzieri was found to be 4.10, 2.20, 4.50 and 6.40 days respectively when they were fed with A. dispersus. The predator had completed its larval development in 17.20 days on the whitefly. The number of whitefly nymphs consumed during first, second, third and fourth larval instar of C. montrouzieri averaged to 23.50, 47.85, 74.60 and 149.80 respectively. The predator consumed a total of 290.75 nymphs during the entire larval development. The predator is being evaluated for its potential in the suppression of the spiralling whitefly under field conditions.

Marotta, S. & Franco, J.C. 2001 (1999). Is the genus Lusitanococcus Neves a junior synonym of Cucullococcus Ferris (Hemiptera: Coccoidea: Pseudococcidae)? Entomologica 33(1999): 127-131. [MarottFr2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The genus Lusitanococcus was established by Neves in 1954 to include arrabidensis, a new species of mealybug collected in Portugal on Erica arborea and E. lusitanica. Later, on the basis of the original illustration and description, some workers considered that Lusitanococcus was a subjective synonym of Cucullococcus Ferris, 1941, while others considered it a valid genus. Specimens were collected in the topotypic locality in Portugal off the same species of host plant and compared with the type specimens of Lusitanococcus arrabidensis Neves and with C. vaccinii Ferris, the type species of Cucullococcus. From this study, we have concluded that: (i) the genus Lusitanococcus is a junior subjective synonym of Cucullococcus, and that (ii) arrabidensis is a valid species. Some points with regard to generic characters of Cucullococcus are discussed.

Marotta, S. & Tranfaglia, A. 2001 (1999). A new morphological structure on Kermes vermilio (Planchon) (Hemiptera: Coccoidea: Kermesidae). Entomologica 33(1999): 133-138. [MarottTr2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Kermes vermilio (Planchon) is a common scale insect living on Quercus ilex L. in urban environments in Southern Italy. Morphological studies on the nymphs and prepupae showed the presence of a pair of membranous ventral frontal lobes, situated just antero-medially to each basal antennal segment. Similar lobes are only currently known in the family Eriococcidae. These structures are therefore here recorded in the Kermesidae for the first time, although it is thought probable that they are present on other species of the family. Their function is unknown.

Maschwitz, U., Dorow, W.H.O., Schellerich-Kaaden, A.L., Buschinger, A. & Azarae, H.I. 2000. Cataulacus muticus Emery 1889 a new case of a Southeast Asian arboreal ant, non-mutualistically specialized on giant bamboo (Insecta, Hymenoptera, Formicidae, Myrmicinae). Senckenbergiana Biologica 80(1-2): 165-173. [MaschwDoSc2000]

Notes: Cataulacus muticus Emery 1889 was found frequently in the Gombak Valley on the Malay Peninsula nesting exclusively in living culms of giant bamboo, mostly Gigantochloa scortechinii Gamble 1896. Big monogynous colonies were polydomously distributed in internodes of large living culms, whereas young colonies with a few workers were found in thin bamboo side branches of the crown region. The colonies, which could comprise more than 2100 workers, occupied culm internodes often opened by pyralid caterpillars but also by other insects. Most but not all natural colonies tended Kermicus wroughtoni Newstead 1897 (Sternorrhyncha: Pseudococcidae) as endophytic trophobionts in the nest internode. In the newly founded colonies these mealybugs were lacking and also not all internodes of mature colonies contained mealybugs. In field and laboratory experiments it could be proven that workers of C. muticus do not transport mealybugs to newly occupied nest internodes during colony expansion or nest relocation. Further details of the sociobiology of C. muticus are reported: non-trophobiotic nutrition, diurnality and swarming behavior. The ants are able to keep their internodes free from water influx not only by entrance blocking with their heads but also by cooperative drinking and urination of the fluid outside the nest. As the ants mostly tap phloem sap from their host plants via their trophobiotic mealybugs and do neither protect their feeding sites nor attack potential living prey they have to be regarded as parasites of their host plants.

Matile-Ferrero, D., Étienne, J. & Tiego, G. 2000. [The introduction of two important pests for French Guiana: Maconellicoccus hirsutus and Paracoccus marginatus (Hem., Coccoidea, Pseudococcidae). (In French). Bulletin de la Société Entomologique de France 105(5): 485-486. [MatileEtTi2000]

Notes: [Original title: Introduction de deux ravageurs d'importance pour la Guyane française: Maconellicoccus hirsutus et Paracoccus marginatus (Hem., Coccoidea, Pseudococcidae).] Origins, distributions and hosts are given for these two pests.

Maw, H.E.L., Foottit, R.G., Hamilton, K.G.A. & Scudder, G.G.E. 2000. In: , Checklist of the Hemiptera of Canada and Alaska. NRC Research Press, Ottawa, Ontario, Canada. 220 pp. [MawFoHa2000]

Notes: 93 species from 11 families of scales reviewed: Margodidae (Margarodidae), Ortheziidae, Aclerdidae, Asterolecaniidae, Coccidae, Cryptococcidae, Dactylopiidae, Diaspididae, Eriococcidae, Kermesidae and Pseudococcidae. Distributions and common names in are given English and French.

McGee, G.G. 2000. The contribution of beech bark disease-induced mortality to coarse woody debris loads in northern hardwood stands of Adirondack Park, New York, U.S.A. Canadian Journal of Forest Research 30(9): 1453-1462. [McGee2000]

Notes: The objective of this study was to adjust previously published estimates of coarse woody debris (CWD) volume and basal areas in northern hardwood forests to account for elevated CWD inputs due to beech bark disease (a disease complex of the scale insect, Cryptococcus fagisuga Lindinger, and a fungus, Nectria spp., on American beech, Fagus grandifolia Ehrh.). Ratios of snags/live tree densities and downed CWD volume/live tree basal area were compared between beech and the codominant, shade-tolerant sugar maple (Acer saccharum Marsh.). The differences between the beech and the sugar maple ratios were used to define the elevated mortality from beech bark disease.

McKenzie, E.H.C., Buchanan, P.K. & Johnston, P.R. 2000. Checklist of fungi on Nothofagus species in New Zealand. New Zealand Journal of Botany 38: 635-720. [McKenzBuJo2000]

Notes: An annotated list is provided of 906 taxa of fungi (including oomycetes and myxomycetes) which have been recorded in New Zealand in close association with the five endemic taxa of Nothofagus (southern beech), as ectomycorrhizal mycobionts, pathogens, or saprobes causing decay of wood and leaves. Nothofagus forests are an important conservation resource, and a vast storehouse for New Zealand fungi; approximately 35% of the known New Zealand agaric mycota are associated with Nothofagus, and 50% of the known polypore species. Sooty mould fungi, growing on honeydew secreted by scale insects, produce conspicuous black growth. Species mentioned as associated with the fungi include Coelostomidia, Crystallotesta fagi and Ultracoelostoma spp.

Mendel, Z., Gross, S., Steinberg, S., Cohen, M. & Blumberg, D. 2001 (1999). Trials for the control of the citrus mealybug in citrus orchards by augmentative release of two encyrtid parasitoids. Entomologica 33(1999): 251-265. [MendelGrSt2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Since the 1980s, the citrus mealybug Planococcus citri (Risso) has become a key pest in Israel, owing to intensive planting of highly susceptible varieties, the introduction of Insect Growth Regulators (which adversely affect coccinellids) and the development of resistance to chlorpyrifos. Management of the mealybug populations in citrus orchards by augmentative releases of parasitoids was investigated between 1993 and 1996 in a series of tests involving the release of 5,000-10,000 Leptomastix dactylopii (Howard) (Encyrtidae) per hectare. Results showed no significant effect on the density of the mealybug on the fruits. In general, the establishment of L. dactylopii was poor and population levels were inferior to those of the naturally occurring Anagyrus pseudococci (Girault) (Encyrtidae). In 1996-1997, augmentative early-spring releases of A. pseudococci at the rates of 10,000-50,000 individuals per hectare markedly increased the population density of A. pseudococci during April and June but had no significant effect on either the mealybug infestation or on fruit damage caused by the pest and its fruit moth associates.

Mestre Novoa, N., Hamon, A.B., Baró Oveido, I. & Reyes Hernandez, M. 2001. [New records of Coccoidea (Homoptera: Sternorrhyncha) for Cuba.] Nuevos registros de Coccoidea (Homoptera: Sternorrhyncha) para Cuba. (In Spanish; Summary In English). Insecta Mundi 15(1): 59-61. [MestreHaBa2001]

Notes: Eriococcus dubius, Parasaissetia nigra and Saissetia nigra are recorded for the first time in Cuba. Geographical distributions, collection site, host plants and comments are given.

Meyer, J.R., Nalepa, C.A. & Devorshak, C. 2001. A new species of anicetus (Hymenoptera: Encyrtidae) parasitizing terrapin scale, Mesolecanium nigrofasciatum (Hemiptera : Coccidae). Florida Entomologist 84(4): 686-690. [MeyerNaDe2001]

Notes: Anicetus carolinensis Meyer (Hymenoptera: Encyrtidae) is proposed as the name of a new species found parasitizing terrapin scale, Mesolecanium nigrofasciatum (Pergande) (Hemiptera: Coccidae), in North Carolina, Both sexes are described and illustrated together with notes on the life history, host range, and reproductive behavior of the species.

Miller, D.R. 2001 (1999). Identification of the pink hibiscus mealybug, Maconellicoccus hirsutus (Green) (Hemiptera: Sternorrhyncha: Pseudococcidae). Insecta Mundi 13(3/4): 189-202. [Miller2001]

Notes: [Publication distributed and postmarked 2001; date printed on issue is 1999.] The pink hibiscus mealybug, Maconellicoccus hirsutus (Green), has spread rapidly in the tropical and subtropical areas of the New World especially throughout the Caribbean Island, and has recently been discovered in California, Mexico, and Belize. All instars of the pink hibiscus mealybug are described and illustrated to facilitate discovery of infestations. Comparisons with other common pest species are provided for most of the 8 instars, and a table is included that distinguishes the pink hibiscus mealybug from other pest species in the field.

Miller, D.R. & Miller, G.L. 2002. Redescription of Paracoccus marginatus Williams and Granara de Willink (Hemiptera: Coccoidea: Pseudococcidae), including descriptions of the immature stages and adult male. Proceedings of the Entomological Society of Washington 104(1): 1-23. [MillerMi2002]

Notes: Paracoccus marginatus Williams and Granara de Willink, commonly known as the papaya mealybug was first discovered in the Caribbean in 1994 and was collected in Florida in 1998. To facilitate implementation of control measures we have prepared descriptions, illustrations, and keys for all stages of this species including first instar, second-instar male and female, third-instar male (prepupa) and female, fourth-instar male (pupa) and female (adult), and fifth-instar male (adult). Comparisons are given between P. marginatus and other species of Paracoccus as well as other commonly encountered mealybug species in Florida and the Caribbean.

Miller, D.R., Ben-Dov, Y. & Gibson, G.A.P. 2001 (1999). Scalenet: a searchable information system on scale insects. Entomologica 33(1999): 37-46. [MillerBeGi2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Systematic information on the scale insects of the world is currently being compiled and synthesized in a database system called BASIS. The information is organized by scale insect family and is searchable at a site on the World Wide Web called ScaleNet (http:\\www.sel.barc.usda.gov\scalenet\scalenet.htm). The site provides general information on scale insects, including sections on economic importance, life histories, distribution and ecology, classification and biographies. A query system provides information on valid names and provides a complete systematic catalogue for any valid genus or species. For a particular taxon, queries will give the following information: all hosts of a scale, distribution of a scale, references for a scale, a checklist of all valid species in a family or genus, biological notes and remarks for a scale. It also will give the scales that occur on a particular host, all systematic references on scales that were published between two dates, all references published by an author, all references with any of five selected words in a title or annotations, a list of all scales described by a particular author, all scales from a particular zoogeographic region, country or country subunit, and the scientific name of any common name of a scale.

Miller, D.R., Williams, D.J. & Hamon, A.B. 2001 (1999). Notes on a new mealybug (Hemiptera: Coccoidea: Pseudococcidae) pest in Florida and the Caribbean: the papaya mealybug, Paracoccus marginatus Williams and Granara de Willink. Insecta Mundi 13(3/4): 179-181. [MillerWiHa2001]

Notes: [Publication distributed and postmarked 2001; date printed on issue is 1999.] Paracoccus marginatus Williams and Granara de Willink, here called the papaya mealybug, was first detected in the United States in Hollywood, Florida in 1998. By the end of 1998 it was found in four localities in the state and has since spread to nine localities in five counties. This mealybug appears to have moved through the Caribbean area since its 1994 detection in the Dominican Republic. The pest is reported to cause serious damage to tropical fruit, especially papaya, and has been detected most frequently, in Florida, on hibiscus. It is now known from Antigua, Belize, the British Virgin Islands, Costa Rica, Guatemala, Mexico, Nevis, Puerto Rico, St. Barthelemy, St. Kitts, St. Martin, and the US Virgin Islands. The species is believed to be native to Mexico and/or Central America. Hosts are listed.

Mishra, Y.D., Sushil, S.N., Bhattacharya, A., Kumar, S. & Mallick, A. 2000. Genetic variations in some productivity linked traits of female lac insects, Kerria spp. (Homoptera: Tachardiidae). Journal of Entomological Research. New Delhi 24(2): 121-124. [MishraSuBh2000]

Notes: Lac insect stocks collected from different localities and host plant species were studied for their performance in respect of five productivity linked female traits, viz., life period, fecundity, dry cell weight, diameter of the cell, and rate of sticklac production during the rainy crop season on Flemingia macrophylla by determining intertrait correlation for the former four traits as well as estimating genetic variance and heritability (in a broad sense) for all the five traits. A highly significant positive correlation between the cell weight and life period, cell weight and diameter of cell, fecundity and diameter of cell, as well as life period and diameter of cell was established. On the other hand, non significant correlation of fecundity with cell weight and life period was recorded.

Mishra, Y.D., Sushil, S.N., Chakrabarty, P.K., Bhattacharya, A. & Kumar, S. 2000. Estimating stick lac productivity of lac on ber, Zizyphus mauritiana Lamk. through regression model. Journal of Entomological Research. New Delhi 24(4): 343-348. [MishraSuCh2000]

Notes: For developing multiple regression model for estimating stick lac productivity of lac insect stocks, a multiple regression analysis was conducted by taking density of settlement, per cent initial mortality, sex ratio, female population, mean live cell weight, resin per female, dry cell weight, density at crop maturity, and life period as independent variables; and sticklac productivity as dependent variable. Out of 51 values of correlation coefficient obtained, 24 revealed highly significant positive and 3 significant negative values while the remaining were non-significant. The multivariate equation derived from only four independent variables i.e. from resin per female, dry cell weight, density at crop maturity, and life period was the most efficient and convenient model for estimating lac productivity to the tune of 97.76% (R super(2)) accuracy.

Miyanoshita, A. & Tatsuki, S. 2001. Role of sex pheromones in reproductive isolation between two host races in Aspidiotus cryptomeriae Kuwana (Homoptera : Diaspididae). Applied Entomology and Zoology. Tokyo 36(no. 2): 199-202. [MiyanoTa2001]

Notes: The reproductive isolation between two host races of Aspidiotus cryptomeriae was examined by comparing intra- and inter-racial responses of males to live virgin females and to sex pheromones in the laboratory. No inter-racial copulation of males to live females was observed for either sex combination. These results indicated that ethological isolation was present between the races. We showed by using a crude extracts test that both host races have female sex pheromones. Although an inter-racial response of males to female extracts was observed it was much weaker than the intra-racial responses, suggesting that the sex pheromone system plays an important role in reproductive isolation between the two races. These results support the hypothesis that each host race is a distinct species.

Mkrtchian, L.P. & Sarkissov, R.N. 2001 (1999). Some characteristics of the development of the sex organs in Margarodid males. Entomologica 33(1999): 167-168. [MkrtchSa2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. A comparative study of the male sex organs of three subfamilies of the Margarodidae (namely Margarodinae, Monophlebinae and Coelostomidiinae) has shown that some morpho-functional rearrangements occur in their ontogenesis during the pre-imaginal period. These changes lead to a reduction of the testes and the formation of an evolutionarily new structure, a spermsac filled with semen bundles. During this general process of structural transformation, two types of spermsac construction can be distinguished, which allows the males of these subfamilies to be divided into two groups: 1. A group which has a spermsac with an ejaculatory duct which penetrates the more or less long, sclerotised penis (e.g., as in species of Porphyrophora, Steingelia and Matsucoccus) 2. A second group where the ejaculatory duct opens into the spermsac and the penis is very short and straight. In this group, the duct is turned out during copulation and then lowered into it - as in species of Icerya, Drosicha and Neomargarodes. The inner position of the ducts in the male sex organs of some scale insects (e.g., male Neomargarodes which retain rudiments of the testes) suggests that this is the primitive condition and that the external position of the ejaculatory duct is secondary. In addition, further study of the Margarodidae has shown that the duct has evolved differently in different groups. Thus, in the first group, the penis tube with the ejaculatory duct within it has developed progressively, while in the second group, in which the penis tube is short, the development of the ejaculatory duct seems to have occurred within the spermsac.

Mohammad, Z.K., Ghabbour, M.W. & Tawfik, M.H. 2001 (1999). Population dynamics of Aonidiella orientalis (Newstead) (Coccoidea: Diaspididae) and its parasitoid Habrolepis aspidioti Compere & Annecke (Hymenoptera: Encyrtidae). Entomologica 33(1999): 413-418. [MohammGhTa2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] An experiment was carried out at Giza governorate, Egypt, between Aug. 1995 and Aug. 1997 to study the fluctuation and seasonal abundance of Aonidiella orientalis and its parasitoid Habrolepis aspidioti on Ficus nitida trees. A brief description of the parasitoid is given. The populations of A. orientalis in both years showed three distinct peaks during the summer, whilst the parasitoid probably had four. The effect of such climatic factors as temperature, relative humidity, photoperiod, dew point and wind velocity were also studied and the size of the populations of both the scale and the parasitoid appeared to be correlated with many of them.

Momen, F.M. 2001. Effects of diet on the biology and life tables of the predacious mite Agistemus exsertus (Acari: Stigmaeidae). Acta Phytopathologica et Entomologica Hungarica 36(1-2): 173-178. [Momen2001]

Notes: The biology of A. exsertus was studied using two different insect eggs, Ephestia kuehniella and Parlatoria zizyphus [P. ziziphi] as the food source. The development was faster and reproduction was higher when A. exsertus fed on eggs of E. kuehniella. A total of 97.78 and 75.27 eggs per female were obtained when eggs of E. kuehniella and P. zizyphus were provided, respectively. A diet of E. kuehniella (eggs) provided the greatest female longevity and mean total fecundity which resulted in higher net reproductive rate (Ro) value (61.25), intrinsic rate of natural increase (rm = 0.196) and finite rate of increase (rm = 1.22) per day for A. exsertus. A diet of P. zizyphus (eggs) resulted in close values of T = 21.70; rm = 174; erm = 1.19. The sex ratio of the progeny was strongly female biased (female/total = 0.72 and 0.66) when both eggs of E. kuehniella and P. zizyphus were provided.

Mondal, S. & Ghosh, A.B, 2000. Morphology of different instars of pink sugarcane mealybug, Saccharicoccus sacchari (Cockerell) (Homoptera, Pseudococcidae). Journal of Interacademicia 4(1): 113-130. [MondalGh2000]

Notes: Descriptions with illustrations and a key to the identification of different instars of the pink sugarcane mealybug, Saccharicoccus sacchari (Cockerell) are given. The trilocular, quadrilocular, clusters of quinquelocular and multilocular disc pores secrete waxy dusts, particulate wax, waxy thread-like caudal projections and globular waxy particles, respectively. Oral collar tubular ducts produce waxy cottony fibres and the glands of anal ring secrete waxy tubes.

Mori, N., Pellizzari, G. & Tosi, L. 2001. [First record of the wax scale Ceroplastes ceriferus (Fabricius) (Hemiptera, Coccoidea) in Italy.] Già acclimatato in Italia il coccide Ceroplastes ceriferus (Fabricius) (Hemiptera, Coccoidea). Informatore Fitopatologia 10: 41-43. [MoriPeTo2001]

Notes: This species was recorded in the Verona district on several ornamental plants. It is the first record of this species in Europe. C. ceriferus is probably native to Asia and is now considered nearly cosmopolitan. It is a polyphagous species, has one generation/year and overwinters as an adult female.

Muralidharan, C.M. & Badaya, S.N. 2000. Mealy bug (Maconellicoccus hirsutus) (Pseudococcidae: Hemiptera) out break on herbaceum cotton (Gossypium herbaceum) in Wagad cotton belt of Kachchh. Indian Journal of Agricultural Sciences 70(10): 705-706. [MuraliBa2000]

Myartseva, S.N. 2001. A new species of parasitoid wasp of the genus Encarsia (Hymenoptera: Aphelinidae) from Tamaulipas, Mexico. (In English; Summary In Spanish). Acta Zoologica Mexicana Nueva Serie 82: 13-18. [Myarts2001]

Notes: Encarsia pinella new sp. was reared from the diaspine scale Melanaspis sp. on Pinus spp.

Myartseva, S.N. & Ruíz-Cancino, E. 2000. Annotated checklist of the Aphelinidae (Hymenoptera: Chalcidoidea) of México. (In English; Summary In Spanish). Folia Entomologica Mexicana 109: 7-33. [MyartsRu2000]

Notes: Species listed as hosts include Acutaspis agavis, Antonina graminis, Aonidiella aurantii, A. citrina, Aspidiotus destructor, A. nerii, Ceroplastes sp., Chrysomphalus aonidum, C. dictyospermi, Clavaspis ?subsimilis, Coccus hesperidum, C. viridis, Diaspis echinocacti, D. simmondsiae, Hemiberlesia rapax, Lecanodiaspis rufescens, Lepidosaphes beckii, L. gloverii, Parlatoria pergandii, Pinnaspis strachani, Pseudococcus agavis, Pseudoparlatoria sp., Quadraspidiotus perniciosus, Saissetia oleae, Toumeyella numismaticum and Unaspis citri.

Nakata, T., Sugiura, N. & Takagi, M. 2000. Host searching efficiency of Aphytis yanonensis and Coccobius fulvus (Hymenoptera: Aphelinidae), introduced parasitoids of Unaspis yanonensis (Homoptera: Diaspididae). Science Bulletin of the Faculty of Agriculture, Kyushu University 55(1): 21-25. [NakataSuTa2000]

Notes: Host searching efficiency of female adults of Aphytis yanonensis and Coccobius fulvus, introduced parasitoids of Unaspis yanonensis, was investigated in the laboratory, using mandarin orange (Citrus unshiu) leaf patches (4X4 cm) with host density of 1, 2, 4 and 8 per patch. We recorded host searching and oviposition behavior of individual female parasitoids until they left the patch. In the higher host density A. yanonensis females attacked more hosts than A. fulvus females when comparing the number of host attacked for 30 or 60 minutes after the patch entrance. This was because of the longer oviposition time per host attack in C. fulvus than in A. yanonensis. However, patch resident time of C. fulvus was longer than that of A. yanonensis, and no significant difference was detected in the number of hosts attacked and parasitized when compared to the per patch visit. It was suggested that the host searching efficiency per unit time was higher in A. yanonensis but that the host searching efficiency per patch visit was not different because C. fulvus stayed on the patch longer.

Narendran, T.C., Bijankumar, D. & Das, B. 2000. A new species of the rare genus Austroterobia Girault (Hymenoptera: Pteromalidae: Austroterobiinae) from West Bengal, India. Zoos' Print Journal 15(9): 319-321. [NarendBiDa2000]

Notes: A new species of Pteromalidae, Austroterobia maldica sp. nov., parasitising an undetermined species of Icerya (Coccoidea) associated with mango in West Bengal, India, is described and its affinities are discussed.

Neser, O.C. & Prinsloo, G.L. 2001. Aphytis chionaspis (Hymenoptera : Aphelinidae), a parasitoid introduced to South Africa from Thailand for the control of mango scale, Aulacaspis tubercularis (Hemiptera : Diaspididae). African Entomology 9(2): 199-201. [NeserPr2001]

Neuenschwander, P. 2001. Biological control of the cassava mealybug in Africa: A review. Biological Control 21(3): 214-229. [Neuens2001]

Notes: Among several natural enemies introduced to combat the cassava mealybug, Phenacoccus manihoti (Homoptera: Pseudococcidae), the neotropical parasitoid Apoanagyrus (Epidinocarsis) lopezi (Hymenoptera: Encyrtidae) was the most successful. It established in 26 African countries, causing a satisfactory reduction in the population density of P. manihoti in most farmers' fields. Four conclusions concerning the possible application of the research results to other biological control projects are discussed. (1) Foreign exploration was intensive and should be maintained at this level in other projects, if necessary at the cost of other activities. (2) In the controversy about the amount of research results needed before first releases are made, an understanding of the proper role of quarantine is essential. Whereas quarantine (preferably outside the continent) guarantees nonnoxiousness of natural enemies, only research in the experimental release sites can determine whether a given natural enemy will be efficient. It was confirmed that the released exotic insects did not affect the diversity of the indigenous fauna. Modalities used in this project for the execution of releases, i.e., always on request by and in collaboration with national programs, are recommended for adoption in future projects. (3) Laboratory and field studies established the scientific basis for quantifying the impact of the pest insect and its control by A. lopezi. This was expressed as reduction in pest population levels, yield loss and gain in revenue. Behavior of adult females in searching and choosing hosts was identified as a better predictor of efficiency of a species in the field than life table studies under controlled temperatures. (4) It is concluded that biological control is the basis for integrated pest management. Other interventions, such as cultural methods or the use of resistant varieties, need to be in harmony with biological control because the impact of natural enemies cannot usually be manipulated by the farmer. To achieve sustainability, the aim is to optimize tritrophic interactions among the plant, the phytophagous pest organisms, and their natural enemies, rather than to maximize the effect of a single intervention.

Nickel, O., Chagas, C.M. & Vasconcelos, A.P.A. 2000. [Association of pineapple mealybug wilt with closterovirus-like particles and dsRNA in Bahia, Brazil.] (In Portuguese; Summary In English). Fitopatologia Brasileira 25(2): 200-202. [NickelChVa2000]

Notes: Leaf and root extracts of pineapple Ananas comosus 'Smooth Cayenne' with and without symptoms of pineapple mealybug wilt (PMW) examined under the electron microscope contained closterovirus-like particles ranging in size from 1,000 to 1,500 X 13 nm, with a modal length of 1,200 nm. A few particles of approximately 2,250 nm in length were also observed. The mealybug Dysmicoccus brevipes was found colonizing pineapple. dsRNA in the range of 0.4 to 17.4 Kb was found in leaves of PMW-diseased pineapple 'Smooth Cayenne', wild bromelias (Pseudananas sagenarius and A. bracteatus) and a diseased hybrid of A. comosus ('Perolera' X 'Smooth Cayenne') submitted to analysis by PAGE. These molecules were not found in seedlings. In SDS-PAGE of virus preparations only one major protein was detected, probably the coat protein, with an approximately Mr of 25 kDa. The nature of the relationship between the closterovirus-like particles and PMW is presently under study.

Novartis Agribusiness Chile SA. 2000. (In Spanish). In: , [Integrated pest management programme. Pome fruit orchards.] Programa manejo integrado de plagas. , . 31 pp. [NAC2000]

Notes: Information is presented from an integrated pest management programme in 11 pome fruit orchards in Chile during 1999-2000. Control was based on monitoring of 7 major arthropod pests, including (Quadraspidiotus perniciosus [Diaspidiotus perniciosus], and some diseases, and their natural enemies. An evaluation of the integrated control programme is given and comments from orchard administrators are included.

Nucifora, S. & Watson, G.W. 2001 (1999). Armoured scale insects (Hemiptera: Coccoidea: Diaspididae) new to Sicily: records and observations. Entomologica 33(1999): 207-211. [NucifoWa2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] The central location of Sicily in the Mediterranean basin makes studies of its fauna particularly interesting. Recent collections from the scrub vegetation on the South-East coast of Sicily and the slopes of Mount Etna have produced four new records of Diaspididae for Sicily: Aonidia ?mediterranea (Lindinger), Ferreroaspis hungarica (Vinis), Mercetaspis isis (Hall) and Chionaspis etrusca Leonardi. The presence of the first three species suggests old faunistic links with other regions, mainly eastern Mediterranean. Our findings are presented here in the hope of stimulating more such research.

Obra, G.P.B. & Rejesus, B.M. 2000. Biological studies of Aphytis sp. nr. chrysomphali (Hymenoptera: Aphelinidae). Philippine Entomologist 14(2): 137-147. [ObraRe2000]

Notes: The parasitoid, Aphytis sp. nr. chrysomphali was collected from coconut scale, California red scale and Florida red scale attacking citrus in Aug 1989 from Villaverde, Nueva Viscaya [Philippines]. The biology of the parasitoid on coconut scale, Aspidiotus destructor Signoret reared on squash was studied. The total developmental period of the uniparental (thelytokous) parasitoid from egg deposition to adult emergence ranges from 13-15 days with a mean of 14.18 plus or minus 0.90. The longevity of adult Aphytis sp. nr. chrysomphali fed with honey ranges from 4-22 days with a mean of 8.45 plus or minus 4.12 days whereas starved adults lived only to a maximum of 33 hours. The total fecundity ranges from 9-24 eggs with a mean of 16.44 eggs.

Öncüer, C., Uygun, N., Erkiliç, L.B. & Karsavuran, Y. 2001 (1999). An annotated list of scale insects (Hemiptera: Coccoidea) from Turkey. Entomologica 33(1999): 231. [OncuerUyEr2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Abstract only. A list of the 171 species of Coccoidea from Turkey, belonging to 10 families, is presented. This list is based on the authors' collection data and on bibliographical sources. The most numerous families are Diaspididae (92 species), Coccidae (31 species) and Pseudococcidae (18 species). The remaining families have between 1 and 7 species each. Among the 171 species are 15 species which are considered to be endemic and their distribution was discussed. In our orchard ecosystems, 11 species are of sufficient importance to require control measures, while in our citrus pest management program, 4 species have pest status. In addition to the cultivated plants, non-cultivated plant have many scale insect species because of the zoogeographical situation of Turkey, lying as it does between the Mediterranean, the Irano-Turanian and Euro-Siberian subregions of Palaearctic. This list is unlikely to be complete and further studies are needed.

Oetting, R.D. 2001. Mexican mealybug -- or is it? Greenhouse Business 7(9): 27-28. [Oettin2001]

Notes: Phenacoccus gossypii, Planococcus citri, Pseudococcus longispinus, P. comstocki, P. maritimus and Rhizoecus falcifer are discussed.

Ozaki, K., Takashima, S. & Suko, O. 2000. Ant predation suppresses populations of the scale insect Aulacaspis marina in natural mangrove forests. Biotropica 32(4): 764-768. [OzakiTaSu2000]

Notes: Previous studies on scale insect-ant interactions have focussed on the mutual relationships between scale insects that produce exudates and attendant ants. These studies may have over-emphasized the prevalence of plus/plus interspecific activity and underestimated the negative effects of ants as predators. The results in this report, however, emphasize that understanding complex scale insect-ant interactions will require a broader survey of the different kinds of interactions between scale insects and ants.

Özgökçe, M.S., Yasar, B. & Karaca, I. 2001 (1999). Life tables of Lepidosaphes ulmi (L.) and Palaeolecanium bituberculatum (Targioni Tozzetti) (Hemiptera: Coccoidea) on apple trees in Van Province, Turkey. Entomologica 33(1999): 317-322. [OzgokcYaKa2001]

Notes: [Special Issue: Proceedings of the ISSIS VIII International Symposium on Scale Insect Studies held at Wye College (U.K.), Aug. 31st - Sept. 6th, 1998.] Apples are the most important cultivated crop in Van Province in Eastern Anatolia Region, Turkey, where Lepidosaphes ulmi (L.) and Palaeolecanium bituberculatum (Targioni Tozzetti) (Hemiptera: Coccoidea) are among the most economically important pest species. Life-table data were collected in the field for these two species in two different apple orchards. The development time for each stage was recorded once a month in the winter and every fortnight during the spring, summer and autumn. In each orchard, both shoots and leaves of five randomly selected trees were sampled and each stage of the two scale species was counted. The life-table