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Anastrepha maculata Norrbom

Recognition
Anastrepha maculata belongs to the daciformis species group, which differs from other species of Anastrepha in having the basal third of the scutellum brown, the male phallus extremely short (less than 0.30 mm long) and with the glans absent (as in the dentata group), and with the female eversible membrane dorsobasally with a single, medially interrupted row of very stout scales. It differs from other species of the daciformis group except A. stonei in having the distal arm of the V-band present. It differs from A. stonei as indicated in the recognition section for that species, and by the following characters: C-band covering cell r₂₊₃ to beyond level of R-M; spot in cell dm (middle part of S-band) not extended anteriorly beyond vein M; frons, except for small brown spot on ocellar tubercle, and gena entirely yellow; scutellum with 3 dark brown spots (1 medial) between yellow basal and white apical areas; scutum entirely microtrichose (nonmicrotrichose except lateral margin in A. stonei); posterior half of anatergite dark brown (white except posteroventral corner in A. stonei); and subscutellum and mediotergite entirely dark brown, or latter sometimes dark orange with 3 dark brown stripes (in A. stonei, subscutellum orange, often with medial dark brown spot; mediotergite entirely orange). The female terminalia are relatively short compared to the other 3 Antillean species of the daciformis species group.

Description
Body predominantly yellow to orange.

Head: Yellow except ocellar tubercle usually brown. Posterior orbital seta usually absent (weakly present on one side in 1m paratype).

Thorax: Scutum usually with dark brown spot above postalar seta, sometimes with dark brown spot mesal to intra-alar seta and/or unpaired dark brown spot posterior to acrostichal setae. Medial pale stripe bilobed posteriorly, lobes stout, with distinct anterolateral corner, extended to dorsocentral seta. Presutural dorsocentral pale stripe connected anteriorly to pale area on postpronotal lobe and connected to or narrowly separated from pale sublateral stripe. Presutural lateral pale stripe poorly differentiated, appears to be complete, including posterior part of notopleuron. Scutellum with 3 dark brown spots, one medial and pair on margin between yellow basal and white apical areas. Pleuron with following dark brown spots: two dorsal spots on anepisternum; anterodorsal and posteromedial spots on katepisternum; medial spot on anepimeron; and posterior half of laterotergite. Subscutellum and mediotergite entirely dark brown, or latter sometimes dark orange with 3 dark brown stripes. Scutum and notopleuron entirely microtrichose; scutellum microtrichose basally, nonmicrotrichose on dark brown spots and apical white area.

Wing: Bands yellow, orange and brown. C- and S-bands completely fused along costa, forming uninterrupted marginal band; cell R₁ without hyaline marginal area. C-band covering cell r₂₊₃ to beyond level of r-m. S-band interrupted in medial cells and along vein Cu₁; basally with broad extension across vein Cu₂+A₁ almost to posterior wing margin; middle part not extended anteriorly beyond vein M, but often narrowly connected to V-band; subapically relatively broad, with large hyaline area in cell r₂₊₃ but well separated from vein R₂₊₃; apically of medium breadth, extended to or almost to apex of vein M. V-band complete, not connected to S-band along vein R₄₊₅. Vein M strongly curved apically; M ratio 1.21-1.53.

Wing.

Abdomen: Tergites entirely yellow or orange, without dark brown marks.

Male terminalia: Lateral surstylus very short, barely extended beyond prensisetae. Phallus extremely short, less than 0.30 mm long; glans absent.

Epandrium and surstyli, lateral and posterior.

Female terminalia: Oviscape length 2.58-2.85 mm, 1.03-1.15 times as long as mesonotum. Eversible membrane, on expanded basal part, with most dorsal scales weakly sclerotized, but with one, medially interrupted row of large, strongly sclerotized, hooklike scales. Aculeus slender, no more than 0.03 mm wide, except at base, length 2.33 mm; tip needlelike, nearly circular in cross-section, with a few fine apical serrations, length 0.10 mm, width 0.025 mm.