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Anastrepha anomoiae Norrbom

Recognition
Anastrepha anomoiae is one of three species of the serpentina group with the C- and S-bands broadly fused so that there is no hyaline area in the middle of cell br. It differs from all other known species of Anastrepha in having the proximal arm of the V-band strongly oblique, with the anterior end more proximal than the posterior end, and connected to the S-band near the middle of crossvein R-M. Crossvein DM-Cu, which is covered by this band, is also oblique in this direction or perpendicular to the long axis of the wing. The posterior end of the V-band does not extend basally along the posterior wing margin. The extremely long aculeus tip (0.86 mm long), which tapers slightly subbasally and is bluntly rounded distally, is also distinctive. The very slender distal section of the S-band, and the dark colors of the wing pattern and the darker areas of the body are additional useful diagnostic characters.

Description
Body largely dark orange to dark red-brown with yellow markings. Setae red-brown to dark brown.

Head: Mostly yellow. Frons, face and gena orange to reddish brown in holotype (possibly discoloration). Ocellar tubercle brown. Occiput in holotype with paired small brown spot dorsally and larger spot ventrally, extending to postgena. Facial carina, in profile, concave. 4-5 frontal setae; 2 orbital setae, posterior seta well developed. Ocellar seta weak. Antenna extended 0.75-0.85 distance to lower facial margin.

Thorax: Mostly orangebrown to red-brown with following areas yellow and distinctly contrasting: postpronotal lobe; single medial and paired sublateral vittae on scutum, the slender medial vitta extended nearly full length of scutum, broadened posteriorly, but extended laterally only slightly beyond level of acrostichal seta (at most half distance to level of dorsocentral seta); sublateral vitta extended from transverse suture to or almost to posterior margin, including intra-alar seta; scutellum except extreme base (brown area well separated from basal seta); dorsal margin of anepisternum; most of greater ampulla; large dorsal spot on katepisternum; katepimeron; and most of anatergite and katatergite. Mesonotal darker areas mostly dark orange, but anterolateral corner, lateral margin of presutural area, margin along mesal side of lateral vitta, somewhat U-shaped posterior area, postsutural lateral areas, and notopleuron dark brown; without orange vitta on dorsocentral line. Darker area of anepisternum about half to entirely brown. Anepimeron mostly dark brown. Darker areas of katepisternum, meron and katatergite dark orange or dark brown. Subscutellum and mediotergite dark orange to dark brown. Mesonotum 3.49-4.00 mm long. Scutum microtrichose except for broad medial anterior area extended 1/3-2/3 distance to transverse suture; setulae mostly brown, but yellow on medial stripe and in broad presutural area. Katepisternal seta weak, no longer than postocellar seta, yellowish to pale brown.

Wing: Length 7.5-9.0 mm. Vein M moderately curved apically; section between BM-Cu and R-M 2.4-2.9 times as long as section between R-M and DM-Cu; section between R-M and DM-Cu 0.63-0.69 times as long as DM-Cu. Crossvein DM-Cu perpendicular to long axis of wing or oblique, with anterior end more proximal than posterior end. Pattern mostly dark brown. Basally with large infuscated area formed by fusion of C-band and base of S-band, which are separated only by hyaline to yellowish area covering much of cell bm; basal area mostly brown, but most of cells bc and c and part of cell bcu yellowish to subhyaline, small basal spot in cell dm and small area in cells sc and r1 posterior to bend in vein sc yellow; cell r2+3 basal to R-M and all of cell br brown. Remainder of S-band brown; distal section very slender, at apex of vein R2+3 0.27-0.33 times width of cell r2+3; closely following vein R2+3 so that hyaline spot in cell r1 is elongate along costa; not extended to apex of vein M. Hyaline spot in cell r1 extended to vein R4+5; its apex aligned with R-M. V-band with distal arm absent; proximal arm very slender, strongly oblique and connected to S-band in cell r4+5 near middle of crossvein R-M; not extended basally along posterior wing margin.

Wing.

Abdomen: Orange with yellow and dark brown bands. Syntergite 1+2 with medial to subapical brown band, broader in male; posterior margin yellow, narrowing laterally. Tergites 3 and 4 with broad brown band, narrowed medially; posterior margin yellow, narrowing laterally. Male tergite 5 orange.

Male terminalia: Dorsal posterior margin of epandrium convex with slight medial angle. Lateral surstylus moderately long; in lateral view very slightly curved; in posterior view with moderate basolateral lobe, main part strongly tapered, slender apically. Proctiger with lateral fold separating sclerotized areas. Phallus 4.58 mm long; 1.31 times as long as mesonotum. Glans 0.65 mm long; acrophallus relatively stout.

Epandrium and surstyli, lateral and posterior.

Female terminalia: Oviscape of holotype damaged, but probably more than 3.7 mm long. Eversible membrane with 60-70 large, hook-like dorsobasal scales in triangular pattern. Aculeus 3.66 mm long; base expanded; tip 0.86 mm long, 0.15 mm wide, non-serrate except for very fine serrations on distal fifth, tapered rapidly subbasally to 0.12 mm wide, then parallel sided, extreme apex broad and bluntly rounded. Spermathecae not examined.

Aculeus tip, ventral; extreme apex, enlarged.