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Anastrepha pseudanomala Norrbom

Recognition
Anastrepha pseudanomala is one of the species of the serpentina group with a hyaline area in cell br posterior to the pterostigma that is not extended to vein R4+5. Of the other species with this character, A. pseudanomala is most likely to be confused with A. normalis Norrbom or A. anomala Stone. It differs from both species in having a slightly longer aculeus tip (more than 0.35 mm long) that is broader than in A. anomala (0.15 mm vs. no more than 0.12 mm), and more than half serrate, unlike in A. normalis. The orbital plate sometimes has a triangular brown mark that does not occur in the other two species. Anastrepha pseudanomala further differs from A. normalis in having the middle section of the S-band solidly dark brown anterior and distal to crossvein R-M.

Description
Body largely dark orange to dark red-brown with yellow markings. Setae dark brown.

Head: Mostly yellow. Ocellar tubercle brown. Orbital plate with paired, faint to moderate brown, triangular spot. Facial carina, in profile, concave. 3-4 frontal setae; 1-2 orbital setae, posterior seta well developed if present. Ocellar seta weak, short to minute. Antenna extended 0.80-0.85 distance to lower facial margin.

Thorax: Mostly orange to dark red-brown with following areas yellow and distinctly contrasting: postpronotal lobe; single medial and paired sublateral vittae on scutum, the slender medial vitta extended nearly full length of scutum, broadened posteriorly, but extended laterally only slightly beyond level of acrostichal seta (at most one third distance to level of dorsocentral seta); sublateral vitta extended from transverse suture to or almost to posterior margin, including intra-alar seta; scutellum except extreme base (brown area well separated from basal seta); propleuron; dorsal margin and anteroventral corner of anepisternum; all or most of greater ampulla; dorsal margin of katepisternum; katepimeron; and most of anatergite and katatergite. Mesonotal pattern similar to A. normalis; broad area bordering medial vitta, shorter and narrower area bordering mesal margin of sublateral vitta, and large sublateral presutural area orange. Most of darker areas of anepisternum, anepimeron, meron, and katatergite dark brown. Katepisternum mostly orange. Subscutellum and mediotergite dark brown, narrowly dark orange medially. Mesonotum 3.62-3.70 mm long. Scutum microtrichose except for broad presutural bare area extended more than 2/3 distance to transverse suture; setulae mostly yellow, brown on parts of darker areas. Katepisternal seta weak, much shorter than postocellar seta, yellowish.

Wing: Length 7.57-7.74 mm. Vein M strongly curved apically; section between BM-Cu and R-M 2.25-2.32 times as long as section between R-M and DM-Cu; section between R-M and DM-Cu 0.73-0.75 times as long as DM-Cu. Crossvein DM-Cu oblique, with anterior end more distal than posterior end. Pattern mostly dark brown. C-band and S-band broadly connected in cells r2+3 and br, but separated basally by hyaline area in posterior half of br aligned with pterostigma, hyaline basal sixth of cell dm, and hyaline to yellowish area covering all of cell bm. C-band yellowish to subhyaline in cell bc and posterior half to 2/3 of cell c; and with large yellow area in base of cell sc and cells r1 and r2+3 posterior to pterostigma, extending distally to level of apex of vein R1. S-band with large yellow area in cell dm extending slightly into cell br and nearly touching crossvein R-M; rest of band dark brown, including areas distal to and anterior to R-M; distal section slightly narrowed, at apex of vein R2+3 0.47-0.49 times width of cell r2+3; not extended to apex of vein M. Hyaline spot in cell r1 nearly triangular, interrupted at or absent posterior to vein R2+3 and not extended to vein R4+5; its apex aligned slightly basal to R-M. V-band with distal arm absent; proximal arm separated from S-band, extended narrowly to vein R4+5; extended basally along posterior wing margin almost to vein A1+Cu2 but not connected to extension from base of S-band.

Wing.

Abdomen: Orange with yellow and brown markings. Syntergite 1+2 with band, brown laterally, orange medially; posterior margin broadly yellow, narrowing laterally. Tergite 3 and sometimes tergite 4 with basal bands, broad and brown laterally, paler and narrower medially; posterior margin narrowly yellow on medial 3/4. Tergites 5-6 orange.

Male terminalia: unknown.

Female terminalia (based on dissected terminalia of paratype and exposed aculeus tip of holotype): Oviscape 4.12-4.28 mm long, 1.11-1.18 times as long as mesonotum. Eversible membrane with 35-40 large, hook-like dorsobasal scales in triangular pattern. Aculeus 4.16 mm long; tip 0.37-0.40 mm long, 0.15 mm wide, parallel-sided basally, apical 0.62-0.63 gradually tapered and very finely serrate. Spermathecae ovoid.

Aculeus tip, ventral.